BIBLIOGRAPHY AND LIST OF ISOETES SPECIES WITH COMPLETE SYNONYMY,

INCLUDING FOSSILS

by Michel Desfayes

Updated January 2008

 

 

Abraham, A. & C.A. Ninan, 1958. Cytology of Isoetes. Current Sci. 27: 60-61.

Allen, B.M. 1975. A note on the distribution of Isoetes in the Cadiz Province, Spain. Fern Gaz. (U.K.) 11(2-3): 163-164.


Allorge, P. 1932. L'Isoetes lacustris L. dans la Chaîne Cantabrique. Cavanillesia 5: 28-30.

Allorge, V. & P. Allorge, 1941. Plantes rares ou intéressantes du nw de l'Espagne. Bull. Soc. Bot. France 88: 226-254.

Alonso, Paz, E. 1989. Notas sobre plantas nuevas o interesantes para la flora Uruguaya: 1. Comun. Bot. Mus. Hist. Nat. Montevideo 5 (91): 1-4. - Isoetes pp. 2-3

Alston, A.H.G. 1945. A new species of Isoëtes from New Guinea. Botanical results of the Richard Archbold expeditions. J. Arnold Arbor. 26: 180.

Alston, A.H.G. 1956. New African ferns. Bol. Soc. Broter. Ser. 2, 30: 5-27.

Alston, A.H.G. 1982. Isoetaceae: 1. In C.G.G.J. van Steenis & R.E. Holtum, eds. Flora Malesiana, series 2. Pteridophytes, volume 1: 62-64. The Hague, Martinus Nijhoff, Dr. W. Junk Publ. 62-64. - illus., chrom. nos., key.

Amigo, J.-J. 1996. Flore et végétation de la partie orientale des Pyrénées. Ptéridophytes, Isoétacées. Naturalia Ruscinonensia, Revue de la Soc. d'Histoire naturelle de Perpignan et des Pyrénées-Orientales, sér. Générale, sect. Botanique, fasc. 6: 1-52.

Amstutz, E., 1957. Stylites, a new genus of Isoetaceae. Ann. Missouri Bot. Garden 44: 121-123.

Andreis, C. & G. Rodondi, 1986, publ. 1987. Alcune stazioni di Isoetes echinospora Dur. nel Bresciano e osservazioni al SEM delle spore delle Isoetes della flora Italica. Natura Bresciana No. 23: 119-130. - illus., maps.

Anthony, N.C., & E.A., 1982. New species and new combinations in African Pteridophyta and Orchidaceae. Contr. Bolus Herb. 10: 143-164.

Anthony, N.C & E.A. Schelpe, 1985. Two new taxa and a new combination in southern African Pteridophyta. Bothalia, 15 (3/4): 554-555 - 1 sp. nov.; 1 var. nov.; 1 comb. et stat. nov.

Arreguin-Sánchez, M., 1986. Nuevos registros y taxa interesantes de pteridofitas del Valle de Mexico (Isoetaceae, Psilotaceae y Selaginellaceae). Phytologia 59(7): 451-453.

Ash, S. & K.B. Pigg, 1991. A new Jurassic Isoetites (Isoetales) from the Wallowa Terrane in Hells Canyon Oregon and Idaho. Amer. J. Bot. 78 (12): 1636-1642.

Aulio, K., 1985. Biomass and chlorophyll contents of Isoetes lacustris as related to water depth in the Lake Pyhäjärvi, SW Finland. Aqua Fennica 15:127-131.

Badré, F. & R. Deschatres, 1979. Les Ptéridophytes de la France, liste commentée des espèces (taxinomie, cytologie, écologie et répartition générale). Candeollea 34: 379-457 (Isoetes p. 427-430).

Bajpai, U. & H.K. Maheshwari, 1984, publ. 1985. SEM studies on the megaspores of Isoetes coromandelina. Phytomorphology, 34(1-4): 226-231. - illus.

Badjpai, A.K., R. Goswami & H.K. Goswami, 2004. Isoetes V: genomic reshuffle [A]: DNA sequences. Bionature 24(1), 25-32.

Baker, J.G. 1880. A synopsis of the species of Isoetes. J. Bot. 18: 65-70, 105-109.

Baldwin, W.K.W. 1933. The organization of the young sporophyte of Isoetes engelmanni, A. Br. Trans. roy. Soc. Canada, ser. 3, sec. V, 27: 11-30.

Barale. G., F. Thévenard, M. Philippe & M. Zarbout, 1998. A new species of the genus Isoetites Münster from the Early Cretaceous of South Tunisia. 5th European Palaeobotanical Palynological Conference Cracow, Abstract p. 11.

Barale, G., 1999. Sur la présence d'une nouvelle espèce d'Isoetites dans la flore du Crétacé inférieur de la région de Tataouine (Sud-Tunisien): implications paléoclimatiques et phylogénétiques. Can. Journ. Bot. 77, 2: 189-196. (I. daharensis nov. sp.)

Baskin, J.M. & C.C. Baskin, 1978. Geographical distribution of Isoetes butleri in the southeastern United States. Amer. Fern J. 68(1): 7-8. - Map.

Bateman, R.M. 1992. Morphogenetic reconstruction, palaeoecology, and phylogeny of Oxroadia gracilis Alvin emend., and O. conferta sp. nov.: anatomically preserved lycopods from Oxroad Bay, SE Scotland. Palaeontographica, vol. 228: 29.

Benl, G. 1988. The Pteridophyta of Bioko (Fernando Po) (Contributions to a flora of the island). 4: Isoetaceae, Pteridaceae, Acrostichaceae, Dennstaedtiaceae, Hypolepidaceae, Athyriaceae, Thelypteridaceae, Blechnaceae. Acta Bot. Barcin. 38: 3-69.

Bennert, H.W., A. Giers, A. Guldenpenning, A. Herbik & A. Katheder, 1992. Some new observations on the fern flora of the Azores. Fern Gaz. 14: 146-148.

Beri, A. & E. Alonso, 1987. Sobre la presencia del género Isoetes en el Uruguay. Resúmenes de las II Jornadas Uruguayas de Botánica. Depto. de Paleontología, Facultad de Ciencias, Universidad de la República, Montevideo.

Berkutenko, A.N. & N.A. Sazanova, 1992. O nakhodke Isoetes maritima (Isoetaceae) v Magadanskoi oblasti. [On the finding of Isoetes maritima (Isoetaceae) in the Magadan region.] Bot. Zhurn. 77(7): 115-117.

Berthet, P. & M. Lecocq, 1977. Morphologie sporale des espèces françaises du genre Isoetes L. Pollen, Spores 19, No. 3: 329 - 359. - illus.

Berthet, P. & R. Pépin, 1984. Isoetes brochonii Motelay est une bonne espèce. Bull. Soc. Bot. France Lett. Bot., 131(2): 139-145. - illus. (but see Prelli & Bock, 1989).

Bhambie, S. 1957. Studies in pteridophytes. I. The shoot apex of Isoetes coromandelina L. J. Indian bot. Soc. 36: 491-502.

Bhambie, S. 1962a. Morphological and anatomical studies in Isoetes and other related genera. Agra Univ. J. Res. Sci. 11(2): 233-6. [Biol. Abstr. 43 (1963) No. 24862.]

Bhambie, S. 1962b. Studies in pteridophytes. II. A contribution to the anatomy of the axis of Isoetes coromandelina L. and some other species. Proc. Indian Acad. Sci. 56: 56-76.

Bhambie, S. 1963a. Studies in pteridophytes. III. On the structure and development of the leaf and sporophyll of Isoetes coromandelina. Proc. Nat. Inst. Sci. India 29B: 169-90.

Bhambie, S. 1963b. The development, structure and organization of root in Isoetes coromandelina L. Proc. Indian Acad. Sci. 58B: 153-64. [Biol. Abstr. 45 (1964), No. 56811.]

Bhambie, S. 1971. Studies in pteridophytes. VIII. An appraisal of the nature of the rhizomorph in Isoetes. J. Indian bot. Soc. 50(1): 56-62.

Bhardwaja, N. & C.B. Gena, 1992 Cytological studies of Rajasthan species of Isoetes L. In Bhardwaja T.N., Gena C.B., eds. Perspectives in pteridology: present and future (Professor S.S. Bir commemoration volume). Part 2. New Delhi: Today and Tomorrow's, 1992 pp.495-499 - series (Aspects of plant sciences; v.14) ISBN 817019413X. Chromosome numbers.

Bhardwaja, T.N. 1984. Three new species of genus Isoetes L. from Rajasthan, India. J. Bombay Nat. Hist. Soc., 81 (1): 165-168 (illus., key).

Bhu, I. & H.K. Goswami, 1990. A new line of chromosomal evolution in Isoetes. Bionature 10(1-2): 45-53.

Blanca, León &  Kenneth R. Young, 1996. Aquatic plants of Peru: diversity, distribution and conservation. Biodiversity and Conservation 5 (10): 1169-1190.

Bociag, K. 1999. New sites for Isoetes echinospora (Isoetaceae) in the Pomeranian lakes (NW Poland). Fragm. Flor. Geobot. 44 (2): 423-427.

Bock, W. 1962. A study on fossil Isoetes. J. Paleontol. 36: 53-59.

Bohra, D.R., B.D. Sharma & R. Singh, 1980. Isoetes in Rajasthan: a study of spores. In Nair, P.K.K., Nagaraj, M., Agashe, S.N., eds. Advances in pollen-spore research: volumes 5-7. New Delhi, Today and Tomorrow's Printers and Publishers.

Boivin, Bernard, 1961. Isoëtes echinospora Durieu in North America. Amer. Fern. J. 51 (2): 83-85.

Bolin, J.F., R.D. Bray, M. Keskin, L.J. Musselman, 2008. The Genus Isoetes L. (Isoetaceae, Lycopyta) in south-western Asia. Turkish J. Bot. 32: 447-457. (I. olympica, I. anatolica, I. duriei, I. subinermis, I. histrix

Boom, B.M. 1979. Spore ornamentation as a useful character in delimiting Isoetes taxa in the southeastern United States. A.S.B.Bull. 26 (2): 75 - key.Boom, B.M., 1980. Instersectional hybrids in Isoetes. Amer. Fern J. 70: 1-4.


Boom, B.M. 1982. Synopsis of Isoetes in the south eastern United States. Castanea, 47 (1): 38-59 - maps, key.

Boom, B.M. & A.M. Evans, 1979. Isoetes butleri in Georgia. Amer. Fern J. 69 (2): 62.

Bory de Saint-Vincent, J.-B., 1844. Sur les Isoëtes et les espèces nouvelles de cette famille découvertes in Algérie. Compte-rendu séances Acad. Sc., Paris 18(26): 1163-1168.

Bosc, L.A.G., 1803. Isoète, Isoëtes. Nouv. Dict. Hist. Nat. 12: 169.

Bose, M.N. & S.K. Roy, 1964. Studies on the upper Gondwana of Kutch - 2. Isoetaceae. Paleobotanist 12: 226-227.

Boston, H.L. & M.S. Adams, 1987. Productivity, growth and photosynthesis of two small `Isoetid' plants, Littorella uniflora and Isoetes macrospora. Journal of Ecology, 75 (2): 333-350.

Boston, H.L., M.S. Adams & T.P. Pienkowski, 1987. Models of the use of root-zone CO2 by selected North American isoetids. Annals of Botany 60: 495-503.

Boutin, C., L. Lesne & A. Thiéry, 1982. Ecologie et typologie de quelquesmares temporaires à isoètes d’une région aride du Maroc occidental. Ecol. Medit. 8: 31–56.  

Braun, A.C.H. 1862a. Anhang über einige auslandische Arten der Gattung Isoetes. Verh. Bot. Vereins Prov. Brandenburg 3: 326-333.

Braun, A.C.H. 1862b. Zwei deutsche Isoetes-Arten nebst Winken zur Aufsuchung derselben. Verh. bot. Vereins Prov. Brandenburg 3-4: 299-333.

Braun, A.C.H. 1864a. Bemerkungen über die Isoetes-Arten der Insel Sardinien. Monatsber. Königl. Akad. Wiss. Berlin, 1863: 554-624.

Braun, A.C.H. 1864b. Les espèces d'Isoetes de l'île de Sardaigne. Ann. Sci. Nat., Bot. sér. V, vol. 2: 306-377. [p. 306-351: discusses the morphology of the following taxa: adspersa, boryana, brachyglossa, capsularis, coromandelina, duriaei, dubia, echinospora, engelmannii, gardneriana, gymnocarpa, histrix, japonica, karstenii, lacustris, lechleri, longissima, malinverniana, riparia, "regulensis" [= tegulensis], setacea, tenuissima, tripus, velata].

Braun, A.C.H. 1868. Über die australischen Arten der Gattung Isoetes. Monatsber. Königl. Akad. Wiss. Berlin 1868: 523-545.

Brautigam, S., C. Li, A. Mudry, G. Miller & E. Prebil, 2001. An analysis of the evolution of the vascular cryptogams, Lycopodophyta, Sphenophyta, and the Pteridophyta, through examination of extinct and extant genera. J. Syst. Biology, Susquehanna University Vol. 8, Number 4.

Briquet, J., 1910. Prodrome de la flore corse. Georg et Cie., Genève, Bâle, Lyon.

Britton, D.M. 1991a. A hybrid Isoetes, I. x harveyi, in northeastern North America. Canad. J. Bot. 69(3): 634-640 (1991) - illus., map.

Britton, D.M., 1991b. Engelmann's quillwort, Isoetes engelmannii, an addition to the  aquatic flora of Canada. Ottawa Field-Naturalists' Club.

Britton, D.M. 1993 Isoetes reticulata R.S. Hill 1987 (Alcheringa 12:158) is an illegitimate name. Amer. Fern J. 83(4): 128

Britton, D.M. & D.F. Brunton, 1989. A new Isoetes hybrid (Isoetes echinospora x riparia) for Canada. Canad. J. Bot. 67 (10): 2995-3002. - illus., maps.

Britton, D.M. & D.F. Brunton, 1991. The spores and affinities of Isoetes taiwanensis (Isoetaceae: Pteridophyta). Fern Gaz. (UK) 14(2): 73-83 - illus., chromosome numbers, reproductive biology.

Britton, D.M. & D.F. Brunton, 1992. Isoetes x jeffreyi, hybr. nov., a new Isoetes (Isoetes macrospora x Isoetes riparia) from Quebec, Canada. Canad. J. Bot. 70(3): 447-452.

Britton, D.M. & D.F. Brunton, 1993. Isoetes x truncata: a newly considered pentaploid hybrid from western North America. Canad. J. Bot. 71(8): 1016-1025 - illus.

Britton, D.M. & D.F. Brunton, 1995. Isoetes x marensis, a new interspecific hybrid from western Canada. Canad. J. Bot. 73(9): 1345-1353 - illus.

Britton, D.M. & D.F. Brunton, 1996a. Isoetes x pseudotruncata, a new triploid hybrid from western Canada and Alaska. Canad. J. Bot. 74(1): 51-59 - illus.

Britton, D.M. & D.F. Brunton, 1996b. Spore morphology and cytology of Isoetes azorica (Pteridophyta Isoetaceae) and its affinity with North America. Fern Gaz. (UK) 15(4): 113-118 - illus. chromosome numbers, anatomy and morphology, palynology, reproductive biology.

Britton, D.M. & D.F. Brunton, 1996c. Isoetes x pseudotruncata, a new triploid hybrid from western Canada. Canad. J. Bot. 74: 51-59.

Britton, D.M. & J.P. Goltz, 1991. Isoetes prototypus, a new diploid species from eastern Canada. Canad. J. Bot. 69(2): 277-281 - illus., maps, chromosome numbers, keys.

Britton, D.M., D.F. Brunton & S.S. Talbot, 1999. Isoetes in Alaska and the Aleutians. Amer. Fern J. 89(2): 133-141 - illus., maps, chromosome numbers, natomy and morphology, palynology, keys

Brooks, J.H. & R.S. Maples, 1971. A recent find of Isoetes in Louisiana. Amer. Fern J. 61(4): 186.

Brouwera E., R. Bobbink & J.G.M. Roelofs, 2002. Restoration of aquatic macrophyte vegetation in acidified and eutrophied softwater lakes: an overview. Aquatic Botany 73 (4):t, Pages 405-431

Brown, L.E. & R.D. Thomas, 1992. Isoetes engelmanii (Isoetaceae) and Juncus trigonocarpus (Juncaceae) new to Arkansas. Sida 15(1): 162.

Brown, R.C. & B.E. Lemmon, 1984. Plastid apportionment and preprophase microtubule bands in monoplastidic root meristem cells of Isoetes and Selaginella. Protoplasma 123: 95-103.

Brown, R.W. 1939. Some American fossil plants belonging to the Isoetales. J. Wash. Acad. Sci. 29: 261-269.

Brown, R.W. 1958. New occurrences of the fossil quillworts called Isoetites. J. Wash. Acad. Sci. 48: 358-361.

Bruchmann, H. 1874. Wurzeln von Isoetes und Lycopodium. Jenaische Seitschr. Naturwiss. 8: 522.

Brunton, D.F & D.M. Britton, 1991a. Isoetes x hickeyi (Isoetaceae: Pteridophyta) in Canada. Fern Gaz. (UK) 14(1): 17-23 - illus.

Brunton, D.F. & D.M. Britton, 1991b. Isoetes x hickeyi in Canada and the Aleutians. Amer. Fern J. 89(2): 133-141.

Brunton, D.F. & D.M. Britton, 1993. Isoetes prototypus (Isoetaceae) in the United States. Rhodora 95(882): 122-128 - illus.

Brunton, D.F. & D.M. Britton, 1996a. The status, distribution, and identification of Georgia quillwort (Isoetes georgiana: Isoetaceae). Amer. Fern J. 86(4): 105-113 - illus.

Brunton, D.F. & D.M. Britton, 1996b. Taxonomy and distribution of Isoetes valida. Amer. Fern J. 86(1): 16-25 - illus., maps, chromosome numbers.

Brunton, D.F. & D.M. Britton, 1997. Appalachian quillwort (Isoetes appalachiana, sp. nov.; Isoetaceae), a new pteridophyte from the eastern United States. Rhodora 99: 118-133 - illus.

Brunton, D.F. & D.M. Britton, 1998. Isoetes microvela (Isoetaceae), a new quillwort from the coastal plain of the southeastern United States. Rhodora 100: 261-275 - illus. maps, chromosome numbers, anatomy and morphology, palynology, keys.

Brunton, D.F. & D.M. Britton, 1999a. Isoetes x echtuckerii. Maritime quillwort, Isoetes maritima (Isoetaceae), in the Yukon Territory. Canad. Field-Nat. 113(4): 641-645. - maps, chromosome numbers, anatomy and morphology, palynology, reproductive biology.

Brunton, D.F. & D.M. Britton, 1999b. Rush quillwort (Isoetes junciformis, sp. nov.), a new pteridophyte from southern Georgia. Amer. Fern J. 89(3): 187-197. - illus. chromosome numbers, anatomy and morphology.

Brunton, D.F. & D.M. Britton, 2006. Isoetes melanopoda subsp. silvatica (subsp. nov.), a new quillwort (Isoetaceae) from eastern North America. Castanea 71: 15-30.

Brunton, D.F. & D.M. Britton, Taylor, W.C. 1994. Isoetes hyemalis, sp. nov. Isoetaceae: a new quillwort from the southeastern United States. Castanea 59(1): 12-21 - illus. maps, chromosome numbers, anatomy and morphology, palynology

Brunton, D.F. & W.C. Taylor, 1990. Isoetes x brittonii hyb. nov. (Isoetaceae): a naturally occurring hybrid (I. engelmannii x I. riparia) in the eastern United States. Amer. Fern J. 80(3): 82-89 - illus., map.

Brunton, D.F., D.M. Britton & T.F. Wieboldt, 1996. Taxonomy, identity, and status of Isoetes virginica (Isoetaceae). Castanea 61 (2): 145-160 - illus.

Budke, J.M., R.J. Hickey, & K.D. Heafner, 2005. Analysis of morphological and anatomical characteristics of Isoetes using Isoetes tennesseensis. Brittonia 57 (2): 167-182.

Burbanck, M.P. & R.B. Platt, 1964. Granite outcrop communities of the Piedmont Plateau in Georgia. Ecology 45: 292-306.

Burrows, John, E. 1990. Southern African ferns and fern allies. Frandsen Publishers Sandton.

Campbell, D.H. 1891. Contributions to the life history of Isoetes. Ann. Bot. 5: 231-256.

Camus, J.M. et al., 1988. Hybridization and speciation in north temperate Isoetes. BSBI News no.49: 41-42.

Caplen, C.A. & C.R. Werth, 2000a. Isozymes of the Isoetes riparia complex, 1. Genetic variation and relatedness of diploid species. Syst. Bot. 25 (2): 235-259. 2. Ancestry and relationships of polyploids. Syst. Bot. 25 (2): 260-280 - illus.

Carroll, N. & R.D. Thomas, 1981. Isoetes melanopoda in Sabine Parish, Louisiana. Phytologia 48 (3): 274 - 275.

Carvalho e Vasconcellos, J. de, 1968. Pteridófitas de Portugal continental e ilhas adjacentes. Fundação Calouste, Gulbekian, Lisboa, pp. 1-190.

Casper, S.J. & H.-D. Krausch, 1980. Pteridophyta und Anthophyta. In Süsswasserflora von Mitteleuropa. H. Ettl, J. Gerloff & H.Heynig, eds., Bd. 23, Teil 1.

Cavot, A., E. & Y. Lemoigne, 1978. La ligule chez les ptéridophytes fossiles et actuelles. Bull. mens. Soc. linn. Lyon 47, 506-16: 581-94.

Cesca, G. & L. Peruzzi, 2001. Isoetes (Lycophytina, Isoetaceae) with terrestrial habitat in Calabria (Italy). New karyological and taxonomical data. Flora Medit. 11: 303-309 - illus.

Ceska, Adolf & Oldriska, 2001. Isoetes minima A.A. Eaton, an overlooked terrestrial quillwort of the pacific northwest. Bot. Electronic News No. 269.

Chang, H.J. & K.S. Hsu, 1977. (Isoetes taiwanensis De Vol) Quart. J. Chinese Forest. 10 (2): 138 - 142 - illus., map.

CHEN Jin-Ming; Jing-Yuan WANG Jing-Yuan; Liu XING; Yan-Wen ZHANG;  Qing-Feng WANG, 2004. RAPD analysis for genetic diversity of Isoetes sinensis. Biodiversity Science 12 (3): 348-353.

CHEN Jin-Ming; Liu XING; Jing-Yuang WANG ; Robert Gituru WAHITI & Qing-Feng WANG, 2006. Genetic variation within the endangered quillwort Isoëtes hypsophila (Isoetaceae) in China as evidenced by ISSR analysis. Aquatic Botany 82: 89-98.

Chen, Y.Y., Q.G. Ye & H.W. Huang, 2003. Preliminary study on allozyme analysis for Isoetes sinensis Palmer. Journal of Wuhan Botanical Research 21:91-94.(in Chinese with English abstract)

Chen, Y.Y.; Q.G. Ye; Z.Z. Li & H.W. Huang, 2004. Genetic structure of Xiuning population of Isoetes sinensis, a critically endangered species in China. Biodiversity Science 12: 564-571.(in Chinese with English abstract)

Chevalier, A. & P. Monnier, 1946. Une espèce d'Isoetes de l'Afrique Occidentale française. Bull. Mus. nation. Hist. nat., Paris, ser 2, 18, 91-97.

Chiang, S.H.T. 1976. The growth cycle of cambium and the structure of the vascular tissue in the corm of Isoetes taiwanensis. Taiwania 21 (1): 14 - 26 - illus.

Chinnock, R.J. 1993. Notes on Isoetes and Tmesipteris in Victoria. Muelleria 8 (1): 57-60 - illus., keys 2 spp. nov.; 1 subsp. nov.

Chung, YH. & H.K. Choi, 1986. (Isoetes coreana, a new species from Korea.) Korean J. Pl. Taxon. 16 (1): 1-12. (illus., anatomy and morphology).

Cieraad, E. & De Lee, 2006. The New Zealand fossil record of ferns for the past 85 million years. New Zealand Journal of Botany 44: 143–170.

Clute, W. N. 1905. The fern allies of North America north of Mexico. Frederick A. Stockes Co., publ.

Cody, W.J. & D.M. Britton, 1989. Ferns and fern allies of Canada. Publication 1829/E, Research Branch, Agriculture Canada, Ottawa.

Cook, C.D.K. 2004. Aquatic and wetland plants of southern Africa. Backhuys Publishers, Leiden, The Netherlands.

Cox, Paul Alan & R. James Hickey, 1984. Convergent megaspore evolution and Isoetes.  The American Naturalist, 124 (3): 437-441.

Croft, J.R. 1985. Ferns and Fern Allies, in Leach, G.J. & Osborne, P.L. 1985. Freshwater Plants of Papua New Guinea. 33 - 74, f. 6 - 13, pl. 5 - 7.

Croft, J.R., 1980. A taxonomic revision of Isoetes L. in Papuasia. Blumea 26: 177-190, f. 1, pl. 1 -21.Illustr., map, key.

Dansereau, P. 1961. Etudes macronésiennes. I. Géographie des cryptogames vasculaires. Agronomia Lusitana 25: 152-181.

Darshan, D. & G.I. Srivastava, 1962. The genus Isoetes in India. Proc. Nat. Inst. Sci. India, vol. 28, part B: 242-280 (with key to species).

De Camp J.D, D.A. Stetler, A.E. DeMaggio, 1994. Expression of sporophytic Storage proteins in the corm of the quillwort (Isoetes echinospora Dur.). Plant Physiol. 106 (4):1395-1402.

Delile, A.R. 1827. Examen de la végétation de l'Isoetes setacea, et exposition de ses caractères. Mém. Mus. Hist. Nat. Paris 14: 100-119.

DeMason, D.A, 1983. The primary thickening meristem: definition and function in monocotyledons. Amer. J. Bot. 70: 955-62.

Dennis, W.M., A.M. Evans & B.E. Wofford, 1979. Disjunct populations of Isoetes macrospora in Southeastern Tennessee. Amer. Fern J. 69 (4): 97 - 99 (1979).

Depoux, M. & A. Pitot, 1965. Etude anatomique de la région ligulaire de la feuille fertile chez Isoetes melanotheca Alston et Isoetes pitotii Alston. Bull. Inst. fr. Afr. noire 27A: 514-50.

Depoux, M. & A. Pitot, 1965. Isoetes pitotii. Bull. Inst. fr. Afr. noire, A, 27: 1-33.

Desvaux, N. A., 1827. Prodrome de la famille des fougères. Ann. Soc. linn., Paris, Mém. 6 (2): 171-212, 6 (3): 213-336.

DeVol, C.E. 1972a. Isoetes found on Taiwan. Taiwania 17 (1): 1-7.

DeVol, C.E. 1972b. A correction for Isoetes taiwanensis DeVol. Taiwania 17 (3): 304-305.

DeVol, C.E. 1975. Flora of Taiwan. (Isoetaceae, I. taiwanensis, ill.)

Dhien, R. 1963. Répartition géographique des Isoetes de la France. Bull. mensuel Soc. Linnéenne Lyon 32: 223-229.

DiMichele, W.A. & R.M. Bateman, 1996, publ. 1997. The rhizomorphic lycopsids: a case-study in paleobotanical classification. Syst. Bot. 21 (4): 535-552. (refers to extant Isoetes).

DiMichele, WA & J.E. Skog, 1992. The Lycopsida: a symposium. Ann. Missouri bot. Gard. 79, 447-9.

Dixon, K.W., J. Kuo & J.S. Pate, 1983. Storage reserves of the seed-like, aestivating organs of geophytes inhabiting granite outcrops in south-western Australia. Aust. J. Bot. 31, 85-103. Corms of Isoetes, stem tubers of Phylloglossum, corms of Stylidium and Philydrella. Minerals, starch grains, protein bodies etc. Also seeds of Stylidium and Philydrella for comparison.

Dodge, R. 1897. A new quillwort. Bot. Gaz. (Crawfordsville) 23: 32-39.

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Goswami, H.K. 1975a. Chromosome studies in natural populations of Isoetes pantii, with heterosporous sporangia. Cytologia 40 (3-4): 543-551.

Goswami, H.K. 1975b. The morphogenetics of mixed sporangia in Isoetes. J. Indian Bot. Soc. 54 (3-4): 210-218 - Illus., chrom. nos.

Goswami, H.K. 1976. A revision of ligule and labium in Isoetes. (Rozwoj, struktura i funkcje ligula i labium u roznych przedstawicieli rodzaju Isoetes). Acta Soc. Bot. Pol. 45 (1 - 2): 69-76 - illus.

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Harms, Vernon L., 1966. Isoetes echinospora var. braunii in interior Alaska.  American Fern Journal 56 (1): 29-32.

Hawes, I., D. Sutherland & D. Hanelt, 2003. The use of pulse amplitude modulated fluorometry to determine fine-scale temporal and spatial variation of in situ photosynthetic activity within an Isoetes-dominated canopy. Aquatic Botany 77 (1): 1-15.

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He, Zican; Hengchang Wang; Jianqiang Li; Qigang Ye & W. Carl Taylor, 2004. Chromosome behavior during meiosis and development of spore mother cells in the Chinese quillwort Isoetes sinensis T. C. Palmer. American Fern Journal, 94 (4): 183-195.

Heafner, K.D. 2002. Noteworthy collections. North Carolina. Castanea 67 (2): 212. Significance. This is the first Isoetes population in North Carolina to conform ecologically, morphologically, and cytogenetically to a published description of I. virginica (sensu Brunton et al. 1996). As such, it is a state record for North Carolina and is the third known extant population of I. virginica in the Southeast. First described by Pfeiffer in 1937, and recircumscribed by Brunton et al. (1996), I. virginica is a tetraploid quillwort of intermittent woodland streams and low, wet wooded areas of the southeastern Mountains and Piedmont.

Heafner, K.D. & R.D. Bray, 2005. Taxonomic reassessment of North American granite outcrop Isoetes Species with emphasis on vegetative morphology and I. piedmontana (Pfeiffer) Reed sensu lato. Castanea, 70 (3): 204-221.

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Henderson, L.F. 1897. A New Isoetes from Idaho. Botanical Gazette 23: 124.

Hickey, R.J. 1981 A new Isoetes from Jamaica. Amer. Fern J. 71 (3): 69 - 74. - illus.

Hickey, R.J. 1984. Chromosome numbers of neotropical Isoetes. Amer. Fern J., 74 (1): 9-13 - illus., chrom. nos.

Hickey, R. J. 1985. Revisionary studies of neotropical Isoetes. Unpublished Ph.D. thesis, The University of Connecticut, Storrs.

Hickey, R.J. 1986a. The early evolutionary and morphological diversity of Isoetes, with descriptions of two new Neotropical species. Syst. Bot. 11: 304-321.

Hickey, R.J. 1986b. Isoetes megaspore surface morphology: nomenclature, variation and systematic importance. Amer. Fern J., 76 (1): 1-16. - illus.

Hickey, R.J. 1986c. On the identity of Isoetes triquetra A. Braun. Taxon 35 (2): 243-246. - illus.

Hickey, R.J. 1988. Isoetes pallida, a new species from Mexico. Amer. Fern J. 78 (1): 35-36 - illus.

Hickey, R.J. 1989. A new species of Isoetes from Territorio Federal Amazonas, Venezuela. Ann. Missouri Bot. Gard. 76 (4): 1160-1162 - illus. Ill., palynology. Isoetes vermiculata sp. nov.

Hickey, R.J. 1990. Notes on neotropical Isoetes L.: 1. A revision of the subgenus Euphyllum, a new subgenus. Ann. Missouri Bot. Gard. 77 (2): 239-245 - illus., map.

Hickey R.J. 1990. Studies of neotropical Isoetes L.: 1. Euphyllum, a new subgenus. Ann. Missouri Bot. Gard. 77. (2): 239-245 - illus., map, keys.

Hickey, R.J. 1994a. Isoetaceae. In: Flora of Peru. Fieldiana, New Series 34:88-97.

Hickey, R.J. 1997a. Genus Isoetes in the New World: an overview. Amer. J. Bot. 84 (6), suppl. 162.

Hickey, R.J. 1997b. Isoetes, pp 254-266 - in Tryon, A.F. and R.A. Moran. The ferns and allied plants of New England. Mass. Audubon Soc. Publ., Lincoln, MA.

Hickey, R. J. & A. R. Smith. 1995. Isoëtaceae. 2: 189–190. In: J. A. Steyermark, P. E. Berry & B. K. Holst (eds.), Flora of the Venezuelan Guayana. Timber Press; Missouri Botanical Garden, Portland; St. Louis.

Hickey, R.J., S.I. Guttman & W.H. Eshbaugh, 1989. Evidence for post-translational modification of triose phosphate isomerase in Isoetes (Isoetaceae). Amer. J. Botany 76: 215- 221.

Hickey, R.J., W.C. Taylor & N.T. Luebke, 1989. The species concept in Pteridophyta with special reference to Isoetes. Amer. Fern J. 79 (2): 78-89 - maps

Hickey, R.J., C. Macluf and W. C. Taylor, 2003. A re-evaluation of Isoetes savatieri Franchet in Argentina and Chile. Amer. Fern J. 93: 126-136.

Hilger, H.H., M. Weigend & W. Frey, 2002. The gametophyte-sporophyte junction in Isoetes boliviensis Weber (Isoetales, Lycopodiophyta). Phyton (Austria) 42 (1): 149-157.

Hill, R.S. 1987. Tertiary Isoetes from Tasmania. Alcheringa 12: 157-162.

Himi, S., R. Sano, T. Nishiyama, T. Tanahashi, M. Kato, K. Ueda & M. Hasebe, 2001. Evolution of MADS-box gene induction by FLO/LFY genes. J. Mol. Evol. 53 (4-5): 387-393.

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Hofstra, D.E. & C.E.C. Gemmill, 1999.  Genetic characterisation of Isoëtes from Lake Omapere.  NIWA Client report (DOC 000229) for Department of Conservation, Northland Conservancy.

Hofstra, D.E., C.E.C. Gemmill & M.D. De Winton, 2006. Preliminary genetic assessment of New Zealand Isoetes and Nitella, using DNA sequencing and RAPDs. Science for Conservation. Science & Technical Publishing, Department of Conservation, Wellington, New Zealand.

Holmer, M., H.S. Jensen, K.K. Christensen, C. Wigand & F.O. Andersen, 1998. Sulfate reduction in lake sediments inhabited by the isoetid macrophytes "Littorella uniflora" and "Isoetes lacustris". Aquatic Botany, 60 (4): 307-324.

Holmes, W.B.K. & S.R. Ash, 1979. Tertiary Isoetes from Tasmania. Alcheringa 12: 157-162.

Holmes, Walter C.,  Ann E. Rushing & Jason R. Singhurst, 2005. Taxonomy and identification of Isoetes (Isoetaceae) in Texas based on megaspore features. Journal of the Plant Resources Center of the University of Texas at Austin, Number 8, December 2005. [Abstract: Scanning electron microscopy examination of Texas specimens of Isoetes revealed differences in megaspore ornamentation patterns of the proximal and distal surfaces that support the recognition of four species of the genus in Texas. These include I. butleri and I. melanopoda, both of widespread occurrence in the central United States, I. lithophila, a central Texas endemic, and I. piedmontana, which we report as new to the state. A key to species based upon megaspore characteristics, distributions, a limited list of exsiccatae, and descriptions and micrographs of megaspores are included.]

HONG, Liu & Qing-Feng WANG, 2003. Aseptic culture of spores in the rare and endangered plant Isoetes sinensis. Tissue culture and rapid propagation of... Plant Physiology Communications 39, No. 5: 219.

HONG, Liu; Qing-Feng WANG & W. Carl Taylor, 2005. Isoetes orientalis (Isoetaceae), a new hexaploid quillwort from China. Novon 15 (1): 164-167.

Hoot, S.B. & W.C. Taylor, 2001. The utility of nuclear ITS, a leafy homolog intron, and chloroplast atpB-rbcL spacer region data in phylogenetic analyses and species delimitation in Isoetes. Amer. Fern J. 91 (3): 166-177.

Hoot, S.B., N.S. Napier & W.C. Taylor. 2004. Revealing unknown or extinct lineages within Isoëtes (Isoëtaceae) using DNA sequences. Am. J. Bot. 91: 899-904.

Hoot, S.B., W.C. Taylor & N.S. Napier. 2006. Phylogeny and biogeography of Isoëtes based on nuclear and chloroplast DNA sequence data. Systematic Botany 31: 449-460.

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HUANG, Q.-F., L. XING & W. Carl Taylor, 2002. Isoetes youngiensis, a new basic diploid quillwort from China. Novon 12 (4): 587-591.

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Abstract. Isoetes L. is a pteridophyte which belongs to Isoetaceae. It has various habitats of submersed, amphibious, terrestrial and alpine. It has two type of spores; megaspore and microspore. Isoetes was divided to four sections by the criteria of megaspore by N. E. Pfeiffer (1922). Isoetes japonica A. BR and I. coreana Chung et Choi was reported in South Korea. Both species are submerged or amphibious. I. coreana (Sect. Cristatae) produces megaspore with irregular, branching crested ornaments on surface. The other species I. japonica (Sect. Cristatae) produces megaspore that has evidently reticulate ornaments. So these two species are distinguishable easily. But reconsideration of the fact whether I. japonica exist or not in South Korea has became to be required. Because a interesting Isoetes material was collected from Island Che-ju. This Isoetes differs from other Isoetes species on the base of many morphological characters. To confirm the identity and phylogenetic relationships among Korean Isoetes we compared ITS sequences for three taxa. (Megaspore, ITS, phylogenetic relationship).

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Karrfalt, E.E. & D.A. Eggert, 1977b. The comparative morphology and development of Isoetes L.: 2. Branching of the base of the corm in I. tuckermanii A.Br.and I. nuttallii A.Br. Bot. Gaz. 138 (3): 357-368. - Illus.

Karrfalt, E.E. & D.A. Eggert, 1978. The comparative morphology and development of Isoetes L. III. The sequence of root initiation in three-and-four-lobed plants of I. tuckermanii A. Br. and I. nuttallii A. Br. Bot. Gaz. 139, 271-83.

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Karrfalt, E.E. 1980. A further comparison of Isoetes roots and stigmarian appendages. Canad. J. Bot. 58 (22): 2318 - 2322 - illus.

Karrfalt, E.E. 1982. Secondary development in the cortex of Isoetes. Bot. Gaz., 143 (4): 439-445 - illus.

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Keeley, J.E. and G. Bowes, 1982. Gas exchange characteristics of the submerged aquatic Crassulacean Acid Metabolism plant, Isoetes howellii. Plant Physiology 70:1455-1458.

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Keeley, J.E., R.P. Mathews, and C.M. Walker, 1983. Diurnal acid metabolism in Isoetes howellii from a temporary pool and a permanent lake. American Journal of Botany 70:854-857.

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Kott, L.S. & D.M. Britton, 1983b. Spore morphology and taxonomy of Isoetes in northeastern North America. Canad. J. Bot., 61 (12): 3140-3163 - illus., maps, chrom. nos., key.

Kott, L.S. & D.M. Britton, 1985. Role of morphological characteristics of leaves and the sporangial region in the taxonomy of Isoetes in northeastern NorthAmerica. Amer. Fern J., 75 (2): 44-55 - illus.

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Luebke, N.T. & J.M. Budke,  2003. Isoëtes tennesseensis (Isoëtaceae), an Octoploid Quillwort from Tennessee. American Fern Journal, 93 (4): 184-190. Abstract. Isoëtes tennesseensis, an octoploid species with a chromosome count of 2n=88, is described. It occurs in the Hiwassee River in Tennessee. Past collections of this species have been misidentified as Isoëtes macrospora (= I. lacustris). Isoëtes tennesseensis differs from I. lacustris in chromosome number, megaspore and microspore morphology and distribution. Speculation on the origin of this new species is presented. Also: Luebke, Neil T., W. Carl Taylor & Mary Ann Polasek, Botany 2002.org, Abstracts.

Luebke, N.T., W.Carl Taylor & M.A. Polasek, 2002. An octoploid species of Isoetes from Tennessee.

http://www.botany2002.org/section11/abstracts/1.shtml

In 1978, populations of Isoëtes were discovered in the Little Tennessee and Hiwassee Rivers of southeastern Tennessee. These populations were reported as disjunct populations of I. macrospora Durieu, a decaploid species (2n=110) of boreal eastern North America. Examination of Isoëtes plants recently collected from the Hiwassee River reveals that these specimens are an undescribed species. Chromosome number and megaspore size and texture distinguish the Hiwassee River plants from I. macrospora. Chromosome counts from root tip squashes of Hiwassee specimens are 2n=88, the first octoploid quillwort reported for North America. Megaspores of Hiwassee River plants are large, ranging in size from 615-950 µm in diameter with an average diameter size of 760 µm. They have a bold rugate-reticulate texture and lack a distinctly ornamented girdle beneath the equatorial ridge. In contrast, megaspores of I. macrospora are smaller, ranging in size from 550-750 µm in diameter with an average of 650 µm. They have a thin cristate-reticulate texture and a prominent papillate to smooth girdle. Comparisons of nuclear ribosomal ITS nucleotide sequences indicate that the Hiwassee River specimens are similar to I. engelmannii A. Braun and I. valida (Engelm.) Clute. In the ITS region, I. engelmannii and I. valida differ from each other by eleven nucleotide substitutions and two 1 bp indels. At seven of these ITS nucleotide sites, the Hiwassee River specimens are like I. engelmannii and at the other four sites they are like I. valida. The Hiwassee River plants share one indel with I. engelmannii and the other indel with I. valida. Isoëtes engelmanii and I. valida, both basic diploids (2n=22), and their allotetraploid, I. appalachiana D.F. Brunton & D.M. Britton (2n=44), are sympatric within the area of the Hiwassee River octoploid.

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Macluf, C. Cecilia, Marta A. Morbelli, Gabriela E. Giudice, 2006. Microspore Morphology of Isoetes Species (Lycophyta) from Southern South America. Part I. Bot. Review 72 (2): 121-134. [Microspores of the 24 species of Isoetes that grow in southern South America were analyzed under a light microscope and scanning electron microscope. The microspores are monolete, elliptic in polar view, 30-40 [micro]m long, and 20-25 [micro]m wide. A background with various characteristics is seen on each surface. A supra-laesural expansion is present. The perispore is ornamented and has a perforated background...]. Part II. TEM analysis of some selected types Bot. Review 72 (2): 135-152 [The microspores of Isoetes escondidensis, I. gardneriana, I. herzogii, I. pedersenii, and I. savatieri were analyzed with transmission and scanning electron microscopy. The selected species were found to be representative of the diversity found in 24 taxa previously studied that grow in southern South America. The sporoderm is similar in the five types and is composed, from the outside to the inside, of perispore, ...]

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Prada C. & C.H. Rolleri, 2005. A new species of Isoetes (Isoetaceae) from Turkey, with a study of microphyll intercellular pectic protuberances and their potential taxonomic value. Botanical Journal of the Linnean Society 147  (2): 213- 228. Abstract. Isoetes anatolica sp. nov. is described from a population growing in seasonal ponds of a mountain near the southern coast of the Black Sea in Bolu, Turkey. It is a robust, amphibious quillwort, characterized by semiterete, carnose microphylls, semicircular foliar section, smooth cuticle, prominent cuticular pegs, stomata, several collenchymatous strands, abundant pectic filaments and connections in the cells of the translacunar diaphragms, incurved alae, well developed ligula, small carnose labium, no velum, well-formed bulliform megaspores and obscurely muriform microspores. Plants were investigated anatomically and a description with additional diagnostic characters is included. Morphological affinities with other species of the genus are discussed. Intercellular pectic protuberances (IPP) were studied in the cells of the translacunar diaphragms of the microphylls of several species of Isoetes including I. anatolica. The IPP were examined to determine if they could provide diagnostic characters. They were detected with TBO and analysed using light and scanning electron microscopy. Types of IPP and species bearing them were as follows: warts in I. adspersa, I. andina, I. boliviensis, I. duriei, I. engelmannii, I. lechleri, I. longissima, I. melanopoda, I. storkii, I. velata subsp. velata, and I. velata subsp. asturicense; warts and filaments in I. brochonii, I. lacustris, and I. setacea, and connections in I. anatolica and I. malinverniana. IPP are lacking in I. boryana, I. echinospora, I. histrix and I. novogranadensis. Combination of type, density, and distribution of IPP promises to be a useful vegetative character in a genus in which diagnostic characters are scarce. 

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Abstract. Phylogenetic trees derived from nuclear ribosomal ITS sequences reveal several well-supported lineages within Isoëtes e.g., an Australasian clade, a southern Africa clade, and a clade containing species from Europe, northern Africa, and western North America. Caribbean species I. cubana and I. jamaicensis and Central American I. panamensis are in a more ancient "Gondwanan" clade that includes I. coromandelina from India. Isoëtes lechleri, I. novo-granadensis, and I. savatieri from South America and I. storkii from Central America form a clade of recently derived and rapidly radiated taxa that also includes most of the species from North America. Morphology, chromosome counts, and DNA sequences indicate that there are undescribed species and interspecific hybrids between the South American members of this clade. Study of Isoëtes collections from the paramos of Ecuador and Venezuela reveal populations of multiple taxa growing together with chromosome counts of 2n=22, 33, 44, 55, 66, 88, and 132, indicating a pattern of reticulate evolution similar to that found in North America. Plants with 2n=33, 55, and 66 produce irregularly formed spores typical of interspecific hybrids. Morphology and chromosome counts were used to construct hypothetical pedigrees of the Andean Isoëtes that will be further tested with ITS and LFY sequence data. Morphology and chromosome counts suggest that Isoëtes killipii, which appears to be an octoploid (2n=88), has crossed with an undescribed tetraploid species (2n=44) to form a hexaploid hybrid (2n=66) that subsequently doubled its chromosome number to form the dodecaploid, I. novo-granadensis (2n=132). Similarly, I. karstenii, a basic diploid (2n=22), has crossed with a tetraploid, possibly I. palmeri, to produce a triploid hybrid (2n=33) and with I. andina, an octoploid, to produce a pentaploid hybrid (2n=55). Morphology, chromosome counts, and DNA sequences provide clues to resolving a phylogeny in Isoëtes.

Taylor, W.C., N.T. Luebke & A.R. Lekschas, 2003. Taxonomic status and evolutionary relationship of Isoetes minima A.A. Eaton (Isoetaceae) based on nuclear ribosomal DNA internal transcribed spacer sequences. Botanical Electronic News, No. 304, March 6, 2003.

Taylor, W.C., A.R. Lekschas, Qing-Feng Wang,  Liu Xing,  N.S. Napier & S.B. Hoot, 2004. Phylogenetic relationships of Isoëtes (Isoëtaceae) in China as revealed by nucleotide sequences of the nuclear ribosomal ITS Region and the second intron of a leafy homolog. American Fern Journal 94 (4): 196-205. Abstract: Isoëtes is an ancient lycopod lineage with a highly conserved morphology that provides few morphological characters to resolve the phylogeny of its species. Species appear to have evolved by divergence and allopolyploidy. The basic diploids I. hypsophila, I. taiwanensis, and I. yunguiensis and the tetraploid I. sinensis occur in China. Analysis of ITS sequences indicates that the Chinese Isoëtes species are part of an Australasian clade including I. brevicula from Western Australia and I. kirkii from New Zealand. Two distinct cloned sequences of the second intron of a leafy homolog were recovered from I. sinensis supporting the hypothesis that I. sinensis is an allotetraploid. One of the I. sinensis cloned sequences was similar to the I. taiwanensis sequence and the other cloned sequence was similar to the I. yunguiensis sequence identifying I. taiwanensis and I. yunguiensis as the likely parents of I. sinensis. Other cloned sequences recovered from I. sinensis were recombined parts of the two distinct sequences. Morphological evidence supporting an allotetraploid origin of I. sinensis is found in its larger microspore size and intermediate megaspore texture compared to I. taiwanensis, and I. yunguiensis.

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Uehara, Koichi, Siro Kurita, Norio Sahashi & Taichi Ohmoto, 1991. Ultrastructural study on microspore wall morphogenesis in Isoetes japonica (Isoetaceae).  American Journal of Botany, Vol. 78, No. 9: 1182-1190.

Umana G., 2001. Limnology of Botos Lake, a tropical crater lake in Costa Rica. Rev Biol Trop. 49 Suppl 2:1-10.

Underwood, L.M. 1888. The distribution of Isoetes. Bot. Gaz. Vol. 13, No. 4: 89-94.

Van De Genachte, E.E., 1996. Conservation genetics of the granite outcrop quillworts Isoetes melanospora and Isoetes tegetiformans. University of Georgia  (dissertation).

Vareschi, V. 1968. Isoetaceae in: T. Lasser (ed.), Helechos, Flora de Venezuela, vol. 1 (1).

Vasudeva, S.M. & S.S. Bir, 1993. Pteridophytic flora of Pachmarhi Hills, central India: 1. General account and families Psilotaceae - Isoetaceae. Indian Fern J. 9(1-2): 153-173 (1992 publ. 1993).

Vega, E., R. Caudales & A. Sanchez Pérez, 1997 New records of pteridophytes for Botswana. Amer. Fern J. 87 (4): 127-130 - Maps.

Verdcourt, B. 2005. Flora of Tropical East Africa. Isoetaceae. Royal Botanic Gardens, Kew 8p. (1 sp. nov., illus., anat and morphology, keys).

Verma, S.C. 1961. Cytology of Isoetes coromandelina. Amer. Fern J. 51: 99-104.

Vines, S.H. On the systematic position of Isoetes. Ann. Bot. 2: 117-123.

Vöge, M., 1997a. Number of leaves per rosette and fertility characters of the quillwort Isoetes lacustris L. in 50 lakes of Europe: A field study. Archiv für Hydrobiologie, 139 (3): 415-431.

Vöge, M., 1997b. Plant size and fertility of  Isoetes lacustris L. in 20 lakes of Scandinavia: A field study. Archiv für Hydrobiologie, 139 (2): 171-185.

Voirin, B., M. Jay & M. Hauteville, 1975. Isoetine, nouvelle flavone isolée de Isoetes delilei et de Isoetes durieui. Phytochemistry, 14 (1):257-259.

Voirin, B. & M. Jay, 1978. Etude chimiosystématique des Lycopodiales, Isoetales, Selaginellales et Psilotales. Biochem. Syst. Ecol., 6: 99-102.

Wagai, S.O., G.K Srivastava & M. Srivastava, 1997. Soil in relation to distribution and growth performance of Isoetes dixitei Shende in the Western Ghats of India. Tropical Ecology 38 (2): 317-322.

Walker, T.G. 1979. The cytogenetics of ferns. Pp. 87-132, in: Dyer, A.F. (ed.). The experimental biology of ferns. London, Academic Press.

WANG, Jing-Yuan; Gituru Robert WAHITI; Qing-Feng WANG, 2005. Ecology and conservation of the endangered quillwort Isoetes sinensis in China.  Journal of Natural History, Volume 39, Number 48, 2005: 4069-4079.

WANG, Qing-Feng; Liu XING; W.Carl Taylor & Zao-Rong HE, 2002, Isoetes yunguiensis (Isoetaceae), a new basic diploid quillwort. Novon 12(4): 587-591 - illus., chromosome numbers, anatomy and morphology, palynology.

Wang W.; M. Tanurdzic, M. Luo; N. Sisneros; H.R. Kim; J.K. Weng; D. Kudrna; C.  Mueller ; K. Arumuganathan; J. Carlson; C. Chapple; C.  de Pamphilis;  D. Mandoli; J. Tomkins; R.A. Wing & J.A. Banks, 2005. Construction of a bacterial artificial chromosome library from the spikemoss Selaginella moellendorffii: a new resource for plant comparative genomics. BMC Plant Biol. 5 (1):10.

WANG, Ziqiang, 1991. Advances on the Permo-Triassic lycopods in North China. I. An Isoetes from the Mid-Triassic in northern Shaanxi Province. Palaeontographica Abt. B Vol. 222: 1-30, 10 pl., 11~fig. keywords: lycopods, Permian-Triassic; China; Isoetes; Shaanxi Province; Permian-Triassic.

WANG, Ziqiang & Lixin WANG, 1982. A new species of the lycopsid Pleuromeia from the early Triassic of Shanxi, China, and its ecology 1982. Palaeontology, 25 (1), 215-225.

Wanntorp, H.-B. 1970. The genus Isoetes in southwestern Africa. Svensk. Bot. Tidskr. 64 (2): 141-157.

Watanabe, M., M. Takamiya, T. Matsusaka & K. Ono, 1996a. Biosystematic studies on the genus Isoetes (Isoetaceae) in Japan: 3. Variability within qualitative and quantitative morphology of spores. J. Pl. Res. 109 (1095): 281-296 - illus. Maps, Chromosome numbers, Anatomy and morphology, palynology, reproductive biology, keys.

Watanabe, M., M. Takamiya, T. Matsusaka & K. Ono, 1996b. Isoetes in South West Africa. Svensk Bot. Tidskr. 64: 141-167. 

Weber, U. 1922. Zur Anatomie und Systematik der Gattung Isoetes L. Hedwigia 63: 219-62.

Weber, U. 1934. Neue südamerikanische Isoetes-Arten. Ber. Deutsche Bot. Ges. 52: 121-125.

Welten, M. 1967. Ein Brachsenkraut, Isoetes setacea Lam., fossil im schweizerischen Mittelland. Bot. Jb. 86: 527-536. (= I. setacea Bosc ex Delile, non Lam.)

West, C. & H. Takeda, 1915. On Isoetes japonica. Trans. Linn. Soc. Lond., ser. 2, Bot. 8: 333-76.

Whatley, J.M. 1974. The behaviour of chloroplasts during cell division of Isoetes lacustris L. New Phytol. 73: 139-42.

Williams, S. 1943. On Isoetes australis S. Williams, a new species from western Australia. Part I. General morphology. Proc. Royal Soc. Edinburgh B62: 1-8.

Wilmanns, O. & H. Rasbach, 1973. Observations on the pteridophytes of São Miguel, Açores. Brit. Fern Gaz. 10: 315-329.

Wolff, H. 1945. Isoëtes lacustre L., eine neue Art für die Gefässpflanzenflora der Schweiz. Verh. schweiz. naturf. Ges. 125: 172-173.

XING, Liu; Y. WANG; Q.F. WANG & Y.H. GUO, 2002. (Chromosome numbers of the Chinese Isoetes and their taxonomical significance). Acta Phytotaxonomica Sinica 40: 351-356. (in Chinese with English abstract)- illus., chromosome numbers, keys.


XING, Liu; X.A. PANG & Q.F. WANG, 2003. Character and variation of chemical properties of the water in the natural habitats of three species of Isoetes in China. Acta Phytoecologica Sinica 27: 510-515.(in Chinese with English abstract)

XING, Liu; Robert Gituru WAHITI & Qing-Feng WANG, 2004. Distribution of basic diploid and polyploid species of Isoetes in East Asia. J. Biogeography 31 (8): 1239-1250. 

XING, Liu; Jing-Yuan WANG & Qing-Feng WANG, 2005. Current status and conservation strategies for Isoetes in China: a case study for the conservation of threatened aquatic plants. Oryx (2005), 39: 335-338 Cambridge University Press. [Abstract. There are four known species of Isoetes (Family Isoetaceae) from China: I. hypsophila, I. sinensis and I. yunguiensis on the mainland, and I. taiwanensis on Taiwan. In a 4-year study we documented the distribution and habitat characteristics of all four species and evaluated their conservation status using IUCN criteria. All four species are facing a high risk of extinction and should be categorized as Critically Endangered on the IUCN Red List. Habitat degradation and loss, water pollution and eutrophication, competitive exclusion, and human disturbance are the main factors causing the decline and extirpation of Isoetes. Only I. taiwanensis is protected in situ in Yangmingshan National Park on Taiwan. None of the species or habitats of Isoetes on mainland China are presently protected, although all species have been designated by the government as State Key Protected Wild Plants. Our results suggest that conservation strategies should be implemented urgently, particularly on the mainland].

Yang, T.Y., S.H.T. Chiang & C.E. DeVol, 1975. Vascular elements in the corm Isoetes taiwanensis. Taiwania 20: 99-106.

Yi, Sun-Young & Masahiro Kato, 2001. Basal meristem and root development in Isoetes asiatica and Isoetes japonica. International Journal of Plant Sciences, 162: 1225-1235.

Yuasa, A. 1933. Studies in the cytology of Pteridophyta. IV. On the spermatozoids of Selaginella, Isoetes and Salvinia. Bot. Mag. Tokyo, 47: 697-709.

Zhang X.C. & L.B. Zhang. 2004. Flora Republicae Popularis Sinicae. Tomus 6(3). Pteridophyta. Huperziaceae, Equisetaceae, Lycopodiaceae, Psilotaceae, Marrattiaceae, Isoetaceae, Cyatheaceae. Beijing: Science Press vii, 313 p. - illus.

Zhu, Shengchao, 2006. Community characteristics and interspecific association of the Songyang population of Isoetes sinensis. Biodiversity Science Vol.14, No.3: 258-264.

Zinda, D.R. 1966. A preliminary report on the comparative morphology of the shoot apex of Isoetes macrospora Dur. and some effects of experimentally applied indole-3-acetic acid and gibberellic acid. J. Minn. Acad. Sci. 33: 107-116. [Biol. Abstr. 48 (1967) No. 71642.]

 

§§§§§

Divisio  LYCOPODIOPHYTA D.H. Scott, Stud. Foss. Pl., ed. 2: 616, 617. 1909.
Common name: clubmosses, quillworts

 

Classis  ISOETOPSIDA J.H. Schaffn., 1910
common name: quillworts, spikemoss

  

Subclassis  ISOETIDAE Reveal, Phytologia 79: 70. 29 Apr 1996.

Order ISOETALES Engler; Syllabus der Pflanzenfamilien, ed. 7, 101 (1912). 1 extant genus, ca. 140 spp. [Isoeteae Barling; Ord. Nat. Plant., p. 16 (1830). Isoetariae Koidz, Acta Phytotax. & Geobot. 7: 11 (1938)]

 

Fossils:

Annalepis brevicystis

Annalepis zeilleri

Illustration: http://www.bioone.org/bioone/?request=display-figures&name=i0002-8444-091-03-0115-f07

Chaloneria cormosa

Isoetites Münster.

Isoetites phyllophila, Isoetites daharensis

Illustration: http://www.bioone.org/bioone/?request=display-figures&name=i0002-8444-091-03-0115-f07

Isoetites horridus (Dawson) Brown

Isoetites serratifolius Bose & Roy

Nathorstiana Richter, 1909.

Nathhorstiana arborea

Illustration: http://www.bio.ilstu.edu/armstrong/syllabi/222book/fig8-14.htm

Oxroadia gracilis Alvin, Oxroadia conforta R.M. Bateman

Paurodentron fraiponti

Pleuromeia sternbergii

Illustration: http://www.bio.ilstu.edu/armstrong/syllabi/222book/fig8-14.htm

Prostigmaria eggertiana

Subordo  ISOETINEAE Engl., Syllabus, ed. 2: 65. Mai 1898.

 

Familia  ISOETACEAE Rchb., Bot. Damen: 309. 1828.
Common name: quillworts  

Subfamilia  ISOETOIDEAE A. Gray, Bot. Text, ed. 5: 502. Jan-Sep 1858.

  

Tribus  ISOETEAE Duby,  Bot. Gall. 1: 543. 12-14 Apr 1828.

 

Genus  ISOETES  L. 1753, Sp. Pl.: 1100 [Calamaria Boehm., Defin. Gen. Pl. (Ludwig) Ed. Boehm. 500. 1760, nom. superfl. Cephalocentron Gennari, Comment. Soc. Crittogam. Ital. 1: 111, 1862. Type Cephaloceraton histrix (Bory) Gennari. Isoëtella Gennari, 1862, Comment. Soc. Crittogam. Ital., 1: 114I). Type I. duriaei, I. lacustris. Stylites Amstutz (1957). Ann. Missouri Bot. Garden 44: 121-123].

 

Note. Lainz, M. 1984. Isoetes, palabra neutra en Latin, esdrujula. An. Jard. Bot. Madrid, 40 (2): 478. (However it has been latinized in the feminine by Linné, and this gender must be maintained as for Orchis).

 

LISTE OF ISOETES SPECIES

 

In the following list I have integrated the Isoetes part of the Checklist of World Ferns on CDROM by Michael Hassler (Germany) and Brian Swale (NZ). http://homepages.caverock.net.nz/~bj/fern/isoetes.htm Distribution data are mostly from Hassler & Swale's CD-rom with some precisions added, especially for South American species.

 

Abbreviations

a.s.l.: above sea level

AWP: Charles D. Stutzenberg, 1999. Aquatic and wetland plants of the western Gulf  Coast. Texas Parks and Wildlife.

C & K: Casper, S. Jost & Heinz-Dieter Krausch, 1981. Pteridophyta und Anthophyta, Band 24,  in H. Ettl, J. Gerloff & H. Heynig, Süsswasserflora von Mitteleuropa. Gustav Fischer Verlag, Stuttgart, New York.

SWUS: Correll, Donovan S. & Helen B. Correll, 1975. Aquatic and wetland plants of southwestern United States. Stanford University Press, Stanford, Calif.

Names in bold type are accepted species or subspecies. Names in italics are synonyms.

 

 

***

Isoetes sp. Crete (Herb. Geneva) = prob. I. delilei

Isoetes sp. Sumatra. Flora Malesiana Bull. 1977, 30: 27-67. Nationaal  Herbarium Nederland (see Flenley & Morley, 1978).

Isoetes sp. Lebanon: Beka'a Valley (2n = 20) (Musselman on line).

Isoetes sp. 2n = 56 (58). Madagascar:  Ambibiroa: route 7, 23 km SW Ambalavao, 800 m east km milestone 23, 1.05.2004  (Herb. Desfayes > Geneva).

 

Isoetes abyssinica Chiovenda, 1933. Atti Soc. Nat. e Mat., Modena 64: 45-47, ill. - Ethiopia, Kenya, Tanzania, Eritrea. See note  sub  I.  welwitschii.

Isoetes acadiensis Kott, 1981. Canad. J. Bot. 59: 2592, 1981 [Isoetes tuckermanii subsp. acadiensis (Kott) Taylor & Luebke] - E Canada (E Newfoundland, New Brunswick, Nova Scotia), NE USA (Massachusetts, Maine, New Hampshire, New York). Map Fl. N Amer.: 70.

Isoëtes adspersa A. Braun in Bory & Durieu,1849, Expl. Sci. Algérie, Atlas: tab. 37, fig. 3] = I. velata subsp. adspersa

Isoetes aequinoctialis Welw. ex A. Braun, 1867, Sitzungsber. Naturf. Freunde, Berlin [Calamaria aequinoctialis (Welw. ex A. Braun) Kuntze, Revisio Generum Plantarum 2: 828 (1891); Calamaria nigritiana (A. Braun) Kuntze, Revisio Generum Plantarum 2: 828 (1891) 1893; Isoetes aequinoctialis Welw., A. Br. in Kuhn, Fil. Afr. 195 (1868); Isoetes erongoensis Wanntorp, Svensk Bot. Tidskr. 64: 150, C 5, t. 2, 3 (1970); Isoetes garnieri Chevalier & Monnier, Bull. Mus, Nation. Hist. Nat. (Paris) 18(1): 91-979 1 fig (1946); Isoetes nigritiana A. Br. ex Kuhn, Filices Africanae 196 (1868)] - Ghana, Mali, Central African Republic, Niger, Nigeria, Zambia, Tanzania, Zimbabwe, Angola, Namibia, Swaziland, N Cape Prov. Ill. COOK, 2004: 33.

Isoetes alata (Pfeiffer) Small = I. flaccida

Isoetes alcalophila Halloy, 1980, Lilloa Rev. bot. 35 (2): 68-75 - Argentina: Tucumán: Laguna Muerta, 4260 m. Ill. microspores MUSSELMAN, 2002.

Isoetes algerica A. Braun, 1848 = I. velata

Isoetes alpina Kirk, Trans. N. Z. Inst. 7: 377, t. 25 (1875) [Calamaria alpina (Kirk) Kuntze, Rev. Gen. Pl. 2: 828 (1891); I. kirkii var. alpina; Isoetes multiangularis Col., Trans. N. Z. Inst. 22: 449 (1890)] - New Zealand (South Isl., North Isl.). See however LARGE & BRAGGINS, 1991: [spore studies] "do not support the existence of two taxa within New Zealand Isoetes (i.e., I. alpina and I. kirkii)". Megaspore variations within the two species overlap".

Isoetes alstonii Reed & Verdcourt, 1965 = I. schweinfurthii

Isoetes amazonica (A.Braun) Baker, 1880 (Calamaria amazonica Kuntze, Revis. Gen. Pl. 2: 828. 1891, I. lacustris sensu Hooker, 1861, Brit. Ferns: 55, non L.) - Brazil: Pará: in vicinibus Santarém =  I. gardneriana

Isoetes amazonica A. Br. = I. gardneriana

Isoetes ambigua = I. setacea subsp. braunii

Isoetes amesii A.A.Eaton = I. riparia

Isoetes anatolica C. Prada & C.H. Rolleri, 2005 (Ill.). 2n = 44 – Turkey: Bolu, S of Abant Golu. Ill. BOLIN et al., 2008.

Isoetes andicola (Amstutz) L.D.Gómez, 1980. Brenesia 18: 4 [Stylites andicola Amstutz, 1957, Ann. Miss. Bot. Gard. 44 (1): 121; Isoetes andicola (Amstutz) H.P.Fuchs, Proc. Kon. Ned. Akad. Wetensch. C 84(2): 177 (1981); Isoetes andicola var. gemmifera (Rauh) L.D. Gómez, Brenesia 18: 4 (1980); Stylites gemmifera Rauh, Sitzungsber. Heidelberg Akad. Wiss. Math. Naturwiss. Kl. 1959: 11 (1959); cf. Reed, C.F. 1953, Bol. Soc. Broteriana 27: 5-72, 28: 5-61] - Andes: Perú: Depto. Lima, prov. Huarochiri supra Casapalca, near Laguna Caprichosa, 4750 m; Puno, Junín. Ill. Tryon, 1982: 831; Fieldiana Bot. 34: 90; MUSSELMAN, 2002 (microspores).

Isoetes andina Spruce ex Hook., British Ferns t. 55, 1861 [?Calamaria triquetra (A. Braun) Kuntze, Revisio Generum Plantarum 2: 828 (1893); Isoetes triquetra A. Br., Verh. Bot. Ver. Brandenb. 4: 332 (1862); ?Isoetes lechleri var. triquetra (A. Braun) L.D. Gómez, Brenesia 18: 5 (1980); Isoetes andina Spruce, Motel. & Vendr., Actes Soc. L. Bord. 36. 325 (1883)] - Venezuela: Mérida: Laguna de Mucubají, 3500 m; Colombia (Laguna de Chisaca, Macizo de Sumapaz), Ecuador, ?S Peru. Ill. Tryon, 1982: 831.

Isoetes appalachiana D.F. Brunton & D.M. Britton, 1997, Rhodora 99: 118-133.

Isoetes argentina Eaton, 1929 - Argentina = I. hieronymii

Isoetes arumiana Fuchs-Eckert, 1959, nom. nud. (Colombia: Cauca: Lagunilla de Las Casitas, 3700 m) = I. fuliginosa

Isoetes asiatica (Makino) Makino, 1914, Bot. Mag. Tokyo 28: 184 = I. setacea subsp. asiatica

Isoetes asturicensis Laínz, 1970, Bol. Inst. Estud. Astur. Supl. Ci. 15: 6, pro subsp. I. boryanae = I. velata subsp. velata

Isoetes atrovirens = I. lacustris

Isoetes attenuata C.R. Marsden & R.J. Chinnock, 1998, Fl. Australia 48: 704 - SE South Australia. Map Marsden, 1979: 334.

Isoetes aucacochensis Fuchs-Eckert, 1981, nom. nud. (Bionature 12: 110, 1992) - Ecuador: Tungurahana: Lake Aucacocha, 3720 m = ? I. ecuadoriensis

Isoetes australis S. Williams, 1943, Proc. Roy. Soc. Edinb. 62B(l). 1-8. 2n = 44 - SW Western Australia. Map Marsden, 1979: 334.

Isoetes azorica Dur. ex Milde, 1867, Fil. Eur., 278 [Calamaria azorica (Dur.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)] - Azores. Cf. Britton D.M. & D.F. Brunton, 1996. Spore morphology and cytology of Isoetes azorica and its affinity with North America. Fern Gazette 15 (4): 113-118 (ill.).

Isoetes baculata R.J. Hickey & H.P. Fuchs, 1986, Syst. Bot. 11(2): 310 - Colombia: Rio Vaupés; Brazil. Ill. Hickey, 1986: 311.

Isoëtes baetica Willk. in Willk. & Lange, 1861, Prodr. Fl. Hisp., 1: 14 Locus class.: In lacunulis limpidis montis Almoráima prope S. Roque = I. velata subsp. velata

Isoetes baptistae Fuchs-Eckert, 1979, nom. nud. (batistae = err.) - Brasil: Rio Grande do Sul: Rio Tainhas ?= I. brasiliensis

Isoetes beringensis = I. maritima

Isoetes biafrana Alston, 1956, Bol. Soc. Brot., ser. 2A, 30: 15 - Cameroon: Lake Oku, Fernando Po: Lago de Moka.

Isoetes bilaspurensis Panigrahi, 1981, Biol. Mems. 6 (2): 135 - India: Madhya Pradesh: Bilaspur = I. coromandelina

Isoetes bischlerae Fuchs-Eckert, 1982, Proc. Kon. Ned. Akad. Wetensch. C 85(1): 113, ill. Locus classicus: Colombia: Boyacá: Páramo de Las Cintas, 3500 m, arroyo, bajo el agua en corrente fuerte - Colombia 3200-358000 m.

Isoetes bolanderi Engelm., 1874, Amer. Nat. 8: 214 [Calamaria bolanderi (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes bolanderi var. parryi Engelm., American Naturalist 8: 214 (1874); Isoetes bolanderi var. sonnei L.F. Hend., Bulletin of the Torrey Botanical Club 27(6): 358 (1900); Isoetes californica Engelm., A Manual of Botany of the Northern United States (ed. 5) 677 (1867); Isoetes parryi Engelm., Motel. & Vendr., Actes Soc. L. Bord. 36: 338 (1883). I. b. var. pygmaea (Engelm.) Clute, Fern Allies, 228, 258 (1905); Calamaria pygmaea (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes bolanderi forma pygmaea (Engelm.) Clute, Fern Bull. 16: 54 (1908); Isoetes pygmaea Engelm., Amer. Nat. 8: 214 (1874)] - W Canada (Alberta, British Columbia), W USA (Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, Utah, Washington, Wyoming, Mexico; altitud. range 1300-3200 m. Map Fl. N Amer.: 73, ill.: 71; Muenscher: 342; SWUS 1: 4, Mason: 30.

Isoetes boliviensis Weber, 1922, Hedwigia 63: 247-248, fig. 14-15 (I. buchtienii Fuchs, 1960, nom. nud.; Isoetes cachukhotaensis Fuchs-Eckert, 1982, nom. nud.; ?= I. feuererii - Bol.: Camacho, 4100 m) - Andes of Peru and Bolivia (Oruro, La Paz: entre Escoma y Hualcapayo; Lago Juriguana near Anilaya, Sorata) 4100-5000 m. Ill.  microspores  MUSSELMAN, 2002.

Isoetes boomii Luebke, 1992, Amer. Fern J. 82: 23 (I. georgiana Luebke, 1992) - USA: Laurens County, Georgia. Ill. Fl. N Amer.: 71.

Isoetes boottii = I. setacea subsp. braunii

Isoetes borboremensis Fuchs, 1960, nom. nud. - Locus classicus: Brasil: Serra Borborema: Lagoa dos Patos = I. luetzelburgii

Isoetes boryana Durieu, 1861 = I. velata subsp. boryana 

Isoetes boryana frigida Gdgr., Fl. Eur. 27: 195 (1891) = I. velata

Isoetes boryana gredensis Gdgr., Fl. Eur. 27: 195 (1891) = I. velata

Isoetes boryana praetervisa = I. boryana

Isoetes boryana var. lereschii = I. velata subsp. velata

Isoetes boryana proles tenuissima (Boreau) Rouy, 1913, Fl. Fr., 14: 477 (comb. illeg.)

Isoetes boyacensis Fuchs-Eckert, 1982, Proc. Kon. Ned. Akad. Wetensch. C Proc. koninkl. nederl. Akad. Wetensch. C 85 (1): 107-109, ill. Locus classicus: Colombia: Santander: Páramo de Almozadero, 3900 m; Cundinamarca, Boyacá, Meta, Nariño. [Isoetes chilesensis Fuchs-Eckert, 1981, nom. nud.] - Colombia: Nariño, Chiles, 3800 m] ?= I. palmeri

Isoetes brachyglossa Braun and = I. coromandelina

Isoetes bradei Herter, 1949, Revista Sudamericana de Botánica 8: 19 (Isoetes gardneriana sensu A.C. Brade) - Brazil: São Paulo. Ill.  microspores  MUSSELMAN, 2002.

Isoetes brasiliensis H.P.Fuchs, 1986, in Reitz, Fl. ilustr. catarinense, Isoetáceas (1): 27, ill. (I. reitzii Fuchs-Eckert, 1979, I. filiformis, I. gonzagensis, ?= I baptistae) - Brazil: Santa Catarina, Rio Grande do Sul, São Leopoldo 10-30 m.s.m. Map FUCHS, 1986: 29, ill.: 23; ill.  microspores  MUSSELMAN, 2002.

Isoetes braunii Durieu = I. setacea subsp. braunii

Isoetes braunii U. Weber, 1922, nom. nud. Locus classicus: Brasil: Serra dos Orgãos, 2238 m (terrestrial, under overhanging rocks) = I. isabelloccensis = I. itatiaiae = I.  organensis

Isoetes brevicula E.R.L.Johnson, 1984, J. Roy. Soc. W. Australia 67(1): 34 - SW Western Australia. Map Marsden, 1979: 335.

Isoetes brittonii = I. englemannii x I. riparia

Isoetes brochonii Motelay, 1892, Actes Soc. Linn. Bordeaux 45: 45 [I. lacustris subsp. brochonii (Motelay) Douin in Bonnier, 1934, Fl. Compl. Fr., 12: 117]. Synonymized with I. lacustris by  R. Prelli & Chr. Bock, 1989, Bull. Soc. bot. France 136: 137-150, but shown to have echinate megaspores by BERTHET & LECOCK, 1977: 335, 357, 358; megaspores similar to those of I. x hickey, but chromosome counts "revealed I. brochonii to be dodecaploid (2n = 12x = 132), double the chromosome number of I. x hickeyi" (Taylor & Hickey, 1992: 620) - France, Spain (E Pyrénées).

Isoetes buchtienii Fuchs, 1960 - Bolivia: dpto. La Paz (nom. nud.) = I. boliviensis 

Isoetes butleri Engelm., 1878, Bot. Gaz. 3: 1 [Calamaria butleri (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes butleri forma immaculata (Engelm.) Clute, Fern Allies, 258 (1905); Isoetes butleri var. immaculata Engelm., Trans. St. Louis Acad. Sci. 4: 388 (1882)] - SC USA: Oklahoma, Missouri, Arkansas, Kansas, N Alabama, C Tennessee, N Illinois, C Texas, Georgia, Kentucky. Map Fl. N. Amer.: 73, Muenscher: 342. Ill. Fl. N Amer.: 71.

Isoetes cachukhotaensis Fuchs-Eckert, 1982, nom. nud. Locus classicus: Bolivia: La Paz: Laguna Cachu Khota, 4140 m =  I. boliviensis. See also Fuchs, 1992: 115.

Isoetes californica = I. bolanderi

Isoetes canadensis (Engelm.) A.A.Eaton ex Maxon = I. riparia

Isoetes capensis Duthie, 1929, Trans. R. Soc. S. Afr. 17: 330. 2n = 22 [Isoetes stephansenii Duthie, Trans. Roy. Soc. So. Afr. 17: 330 (1929), Isoetes capensis var. stephansenii (A.V. Duthie) Schelpe & N.C. Anthony, Bothalia 15(3-4): 555 (1985)] - S Afr.: W Cape Prov. Ill. COOK, 2004: 33.

Isoetes capillacea Bory de Saint-Vincent, 1846 (Oran, Algeria) = I. velata

Isoetes capmanni Engelm., 1883 = I. chapmannii = I. flaccida

Isoetes capsularis Roxb. = male plant of Vallisneria spiralis

Isoetes carchiensis Fuchs-Eckert, 1981, nom. nud. Locus classicus: Ecuador: Carchí, Páramo El Angel, 3450-3500 m.

Isoetes carltaylorii Musselman (Indian River, Canada) = Isoetes acadiensis x Isoetes engelmannii

Isoetes caroli E.R.L. Johnson, J. Roy. Soc. W. Australia 67(1): 34 (1984) - SW Western Australia. Map Marsden, 1979: 333.

Isoetes caroliniana (A.A. Eaton) Luebke, 1992, Amer. Fern J. 82: 26 - Eastern USA: North Carolina, Tennessee, Virginia, West Virginia.

Isoetes castillonii Fuchs-Eckert, 1965, nom. nud. Locus class.: Argentina: Tucumán: Laguna de la Manza, 4200 m (= I. escondidensis)

Isoëtes chaboissaei Hy in Morot, 1894, J. Bot. 8: 97 (Brenne) = I. velata subsp. tenuissima

Isoetes chaetureti E.J. Mendes, 1961, Agron. Lusit. 23: 7 (Estremadura, Portugal) = I. histrix

Isoetes chapmanii Small ex Pfeiff. = I. flaccida

Isoetes chilensis Fuchs-Eckert, 1981, nom. nud.. Locus class.: Argentina: Rio Negro, 800 m - Chile: Coquimbo: Pichindangi, 1000 m, Nahuelbuta 1250 m = I. savatieri

Isoetes chilesensis Fuchs-Eckert, 1981, nom. nud. Locus class.: Colombia: Nariño, Chiles, 3800 m = I. boyacensis

Isoetes chubutiana Hickey, Macluf & Taylor, 2003 – C Chile, C Argentina. Ill.  microspores  MUSSELMAN, 2002.

Isoetes clavata Weber, 1934, Berichte Deut. Bot. Gesellsch. 52: 121 - British Guyana = I. ovata Pfeiffer, 1922 (I. goebelii Christ ined., non Weber, 1922) = prob. I. triangula

Isoetes cleefii Fuchs-Eckert, 1981, Proc. koninkl. nederl. Akad. Wetensch. 84 (2): 177-179, ill. Locus class.: Colombia: Cundinamarca: Páramo de Sumapaz: Laguna Gubernador, 3815 m. = I. palmeri, ?= I. boyacensis

Isoetes colombiana (T.C. Palmer) Fuchs-Eckert, 1981, Proc. Kon. Ned. Akad. Wetensch. C 84(2): 176, "pro var. I. lechleri" [Isoetes lechleri var. colombiana T.C. Palmer Amer. Fern Journ. 19: 18 (1929)] = I. karsteni var.

Isoetes coreana Y.H. Chung & H.K. Choi, 1986, Korean J. Pl. Taxon. 16(1): 2 - Korea I. japonica subsp. sinensis

Isoetes coromandelina L.f., 1781, Suppl. Pl. Syst. Veg. ed. II: 447 (coromandeliana) Indian subcontinent. Cf. Pant & Srivastava (1962), and Fraser-Jenkins (1997). Isoetes coromandelina subsp. coromandelina L. f., Suppl. Pl. Syst. Veg. ed. Il: 447 (1781) [Calamaria coromandelina (L. f.) Kuntze, Revisio Generum Plantarum 2: 828 (1891)-1893; Isoetes bilaspurensis Panigr., Biol. Mem. 6 (2): 135. f. 1-10 (1981), fide Fraser-Jenkins (1997); Isoetes brachyglossa A. Braun, Verh. Bot. ver. Brandenb 4 328 (1862), fide Fraser-Jenkins (1997); Isoetes coromandelina subsp. brachyglossa (A. Braun) Panigr., Biol. Mem. 6 (2): 131 (1981), fide Fraser-Jenkins (1997); Isoetes capsularis Griffith, Posth. Papers, Cryptog. Pl. 572-575, t. 116-118 (1849) (non Roxb.); Isoetes coromandelina var. raipurensis Unni, fide Fraser-Jenkins (1997); Isoetes dixitei Shende, Journ. Univ. Bombay 14: 50 (1945), fide Fraser-Jenkins (1997); Isoetes indica Pant & Srivastava, Proc. Nat. Inst. Sci. India 28 B (3) 246. t. XIV9 f. 16-259 t. XV f. 30-35 t. XIV f. 30-40 text-fig. 4 A-D, 5 A-H, 7 A-M, 12 (1962), fide Fraser-Jenkins (1997); Isoetes mahadevensis G.K. Srivasta, D.D. Pant & P.K. Shukla, Amer. Fern J. 83(4): 106 (1993), fide Fraser-Jenkins (1997); Isoetes mirzapurensis Panigr. & Dixit, Proc. Indian Sci. Congr. pt. III, Abstract, Botany 243 (1965), fide Fraser-Jenkins (1997); Isoetes panchananii Pant & Srivast., Proceedings of the National Institute of Science of India, Part B, Biological Sciences 28: 243, t. 13, 14, f. 1, 2 (1962), fide Fraser-Jenkins (1997); Isoetes panchganiensis G.K. Srivast., D.D.Pant & P.K. Shukla, Amer. Fern J. 83(4): 107 (1993), fide Fraser-Jenkins (1997); Isoetes panchganiensis var. kemangundiensis G.K. Srivast., D.D. Pant & P.K. Shukla, Amer. Fern J. 83(4): 114 (1993), fide Fraser-Jenkins (1997); Isoetes pantii Goswami & Arya, Journ. Indian Bot. Soc. 49: 30 (1970), fide Fraser-Jenkins (1997); Isoetes rajasthanensis Gena & Bhardwaja, J. Bombay Nat. Hist. Soc. 81(1): 165 (1984), fide Fraser-Jenkins (1997); Isoetes reticulata Gena & Bhardwaja, J. Bombay Nat. Hist. Soc. 81(1): 166 (1984), fide Fraser-Jenkins (1997); Isoetes sahyadrii Mahabale, Curr. Sci. 7 (2): 62. f. 1 (1938), fide Fraser-Jenkins (1997); Isoetes sampathkumarnii L. N. Rao, Curr. Sci. 13: 286. f. 1-3 (1944), fide Fraser-Jenkins (1997); Isoetes tuberculata Gena & Bhardwaja, J. Bombay Nat. Hist. Soc. 81(1): 167 (1984), fide Fraser-Jenkins (1997); Isoetes semilocularis J.E. Smith, Rees, Cyclopaedia 19: 150 (1819); Isoetes unilocularis Smith] - India (Tamil Nadu, Kerala, Karnataka, Maharashtra, Gujarat, Madhya Pradesh, Uttar Pradesh, Bihar, Orissa, West Bengal), Sri Lanka, Thailand, Vietnam. Isoetes coromandelina subsp. macrotuberculata C.R. Marsden, Contr. Herb. Austral. 24: 2 (1976) - N Western Australia, Northern Territory, Queensland. Map Marsden, 1979: 332.

Isoetes coromandelina subsp. brachyglossa = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes coromandelina var. raipurensis = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes costaricensis Fuchs-Eckert, 1980, nom. nud., pro syn. I. tryoniana. Locus class.: Costa Rica: Cordillera de Talamanca: Páramo Buena Vista, 3100 m. (= I. tryoniana = I. storkii).

Isoetes cristata C. R. Marsden & Chinnock, 1998, Fl. Australia 48: 704 - Australia: Northern Territory, 10 km south of Jimmy's Creek. Known only from type locality. Map Marsden, 1979: 331.

Isoetes cubana Engelm. ex Baker, 1880, Journal of Botany, British and Foreign 18: 110 [Calamaria cubana (Engelm. ex Baker) Kuntze, Revisio Generum Plantarum 2: 828 (1891)] - Cuba, Mexico (Yucatan), Belize. Ill. Tryon, 1982: 833.

Isoetes cusiyacoensis Fuchs-Eckert, 1981, nom. nud. Locus class.: Colombia: Cauca: Laguna de Cusiyaco, 3200 m. (?= I. fuliginosa)

Isoetes debii P.K.Shukla, G.K Srivastava, S.K. Shukla, 2002 - northeastern India.

Isoetes decipiens = I. velata

Isoëtes delalandei J. Lloyd, 1851, Notes Fl. Ouest France: 25-28 = I. histrix

Isoetes delilei Rothmaler, 1944, Feddes Repert., 54: 72. (2n = 22 - Madrid, Spain) [Isoetes palustris var. setacea F. Mueller, Milde, Fil. Eur. & Atl. 9: 280 (1867) = Isoetes setacea var. delilei Bory, Compt. Rend. Acad. Paris 18: 1165 (1844), and Flora 27: 716 (1844); Isoetes setacea auct. non Lam., 1789] - SE France, Portugal, W Spain [? Corsica (see BRIQUET, 1910: 34), S Sardinia (GLÜCK, 1936) – records uncertain]. Map Cook, 1983: 555. Ill. C & K 1: 52, PRELLI 83; BERTHET & LECOCQ, 1977: 346, 347; ROMERO et al., 2004.

Isoetes dichotoma L.E. Mora & W.Hagemann, 1977, Mutisia 43: 5 - Colombia = I. novogranadensis

Isoetes dispora Hickey, 1994, Fieldiana, Bot. 34: 93 - Peru: Lambayeque, 3300 m. Known from type only.

Isoetes divyadarshani P.D. Shukla, G.K. Srivastava,  S.K. Shukla & P.K. Rajagopal, 2005, Taxon, 54 (1): 109-116. - Peninsular India.

Isoetes dixitei Shende, 1945, J. Univ. Bombay, vol. 14: 50-52 - India: Madhya Pradesh: Panchgani, known only from type locality. Ill. loc.: 51 and in Pant & Srivastava, 1962 = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes dodgei A.A.Eaton = I. setacea x I. riparia

Isoetes dodgei var. robbinsii A.A. Eaton = I. setacea x I. riparia

Isoetes drummondii A. Br., 1863, Monatsber. K. Akad. Wiss. Berlin, 593. Isoetes drummondii subsp. drummondii A. Br., Monatsber. K. Akad. Wiss. Berlin, 593 (1863) [Calamaria drummondii (A. Br.) Kuntze, Rev. Gen Pl. 2: 828 (1891)] - New Guinea, Western Australia, South Australia, Victoria, Tasmania. I. d. subsp. anomala C.R. Marsden & Chinnock, Muelleria 8(1): 58 (1993) - South Australia, Western Australia, Victoria, New South Wales. Maps Marsden, 1979: 332, 333, ill. Aston: 35

Isoetes dubia Gennari, 1861, nom. illeg., non Ludwig, 1859 (Isola Maddalena) = I. velata

Isoetes dubia var. emersa = I. velata

Isoetes dubia var. maculosa = I. velata

Isoetes duriaei balearica = I. duriaei

Isoetes duriaei Bory, 1844 Compt. Rend. Acad. Paris 18: 1166 (1844), 2n = 55 (Calabria - Cesca & Peruzzi), 2n = ca 121-132 [(I. duriei, I. durieui); Calamaria durieui (Bory) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetella duriaei Genn., Comment. Critt. Ital. l(3): 115 (1862); Isoetes duriaei balearica Gdgr., Fl. Eur. 27: 196 (1891); Isoetes duriaei duriaei Bory, Gdgr., Fl. Eur. 27: 196 (1891); Isoetes duriaei felix Gdgr., Fl. Eur. 27: 196 (1891); Isoetes duriaei insularis Gdgr., Fl. Eur. 27: 196 (1891); Isoetes duriaei kabylica Gdgr., Fl. Eur. 27: 196 (1891); Isoetes duriaei leptophylla Gdgr., Fl. Eur. 27: 196 (1891); Isoetes duriaei ligustica DeNot., Kunze, Ind. Fil. Cult., 51 (1850); Isoetes duriaei litorea Gdgr., Fl. Eur. 27: 196 (1891); Isoetes duriaei lusitanica Gdgr., Fl. Eur. 27: 196 (1891); Isoetes duriaei maritima Gdgr., Fl. Eur. 27: 196 (1891); Isoetes duriaei sardoa Gdgr., Fl. Eur. 27: 196 (1891); Isoetes ligustica DeNot., ex A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec. (1863); Isoetes tridentata Dur., ex Kuhn, Fil. Afr. 195 (1867)] - France, Portugal, Spain, Baleares, Italy, Corsica, SardiniaSicily, Malta, Tunisia, Algeria, Morocco, Greece, Turkey: Rizeh (Black Sea). Ill. C & K 1: 52, 59, Prelli: 83; BERTHET & LECOCQ, 1977: 331, 337, 338; ROMERO et al., 2004.

Isoetes duriaei Hook. (Guernesey) = I. histrix

Isoetes dusenii U. Weber, 1920, nom. nud. Locus class.: Brasil: Itatiaia: Serra de Pico Redondo, Rio de Janeiro 2000-2300 m, in streams ?= I. itatiayae, ?=  I. organensis

Isoetes eatoni R. Dodge, Isoetes eatonii forma gravesii (A.A.Eaton) C.F.Reed = I. engelmannii x I. setacea

Isoetes echinospora Durieu de Maisoneuve, 1861, Bull. Soc. Bot. France 8: 164. (2n = 22) = Isoetes setacea, Lam. 1789) [I. muricata Dur., Isoetes echinospora subsp. muricata (Durieu) A. Löve & D. Löve, Isoetes echinospora var. muricata (Durieu) Engelm. in A.Gray. I. hesperina] - ­Eurasia, N Amer. Map Hultén & Fries, 1: 5, Taylor et al., p. 68. Ill. Prelli: 83. - Isoetes echinospora subsp. echinospora - W Palaearctic S to the Pyrenees, N Italy, Bulgaria. Map Hultén & Fries 1: 5; Fl. N Amer.: 68; Muenscher: 343; ill. Fassett: 46; AWP 1: 26; Muenscher: 341; Crow & Hellquist 1: 26. I. echinospora subsp. asiatica (Makino) A.Löve 85 [I. asiatica (Makino) Makino, Bot. Mag. Tokyo 28: 184 (1914); Isoetes echinospora var. asiatica Makino, Bot. Mag. Tokyo 18: 129 (1914), Iversen, Dansk Bot. Ark. 5 (23): 2 (1928); Isoetes muricata Durieu subsp. asiatica (Makino) H.P.Fuchs, Proc. Kon. Ned. Akad. Wetensch. C 85(2): 239 (1982); Isoetes setacea Lam. subsp. asiatica (Makino) Holub; Isoetes echinospora var. savilei B.Boivin] - Mongolia, S Kuril Isl., Sakhalin, Kamchatka, Korea, Japan, ?China. Map Hultén & Fries 1: 5; map and ill. Kadono 8, 10. Isoetes echinospora subsp. braunii [I. braunii Durieu, Isoetes echinospora var. braunii (Durieu) Engelm. in A.Gray; Isoetes tenella var. braunii (Durieu) Rothm.; Isoetes braunii forma astoma (Iversen) M.Braun; Isoetes tenella forma astoma (Iversen) Rothm.; Isoetes echinospora forma astoma Iversen; Isoetes tenella forma iversenii Rothm.; Isoetes braunii forma oligostoma (Iversen) M.Broun; Isoetes tenella forma polystoma Rothm.; Isoetes braunii forma polystoma (Iversen) M.Broun; Isoetes tenella forma oligostoma Rothm.; Isoetes tenella var. oligostoma (Iversen) Rothm.; Isoetes muricata var. braunii (Durieu) C.F.Reed; Isoetes braunii forma robusta (Engelm.) Pfeiffer; Isoetes tenella forma robusta (Engelm.) Rothm. Isoetes echinospora var. brittoni Cockerell; Isoetes echinospora (infrasp. unranked) brittonii Cockerell; Isoetes echinospora forma boottii (Engelm.) Clute; Isoetes echinospora var. boottii Engelm.; Isoetes echinospora subf. angustivelata Iversen; Isoetes echinospora subf. lativelata Iversen; Isoetes tenella subf. lativelata (Iversen) Rothm.; Isoetes tenella forma lativelata (Iversen) Rothm.; Isoetes echinospora var. robusta Engelm.; Isoetes echinospora forma robusta (Engelm.) Clute; Isoetes muricata forma robusta (Engelm.) C.F. Reed; Isoetes muricata var. hesperia C.F. Reed] - Alaska, Canada (Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, Labrador, Nunavut, N.W. Territories, Nova Scotia, Ontario, Prince Edward Isl., Quebec, Saskatchewan, Yukon), NW USA, northeastern USA (Washington, California, Colorado, Connecticut, Idaho, Maine, Massachusetts, Michigan, Minnesota, Rhode Island, Utah, Vermont, Wisconsin). 

Isoetes echinospora subsp. asiatica var. savilei Boivin, 1961, Amer. Fern J. 51: 85 = I. setaceaa subsp. asiatica

Isoetes setacea x I. riparia (I. x eatonii)

Isoetes ecuadoriensis Asplund, 1925, Botaniska Notiser 1925: 357 - Ecuador: prov. Imbabura in Laculo montis Mojanda, alt. 4000 m.s.m. 

Isoetes ekmanii U. Weber, 1922, Hedwigia 63: 251. fig. 26-27 - Argentina: Buenos Aires, Misiones. Ill.  microspores  MUSSELMAN, 2002. (= I. paraguayensis - Misiones, Pilcomayo, President Hayes, dept. Central).

Isoetes elatior F. Muell. & A. Br., 1852, Linnaea 25: 722 [Calamaria elatior (F. Muell.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes tasmanica F. Muell., Dur., Bull. Soc. Bot. Fr. 11: 104 (1864)] - Australia, Tasmania. Map Marsden, 1979: 338.

Isoetes engelmannii A. Br., 1846, Flora 29: 178. 2n = 22 [Calamaria engelmannii (A. Br.) Kuntze. Rev. Gen. Pl. 2: 828 (1891); Isoetes engelmannii var. fontana A.A.Eaton, Fern Bull. 13: 52 (1905); Isoetes engelmannii forma fontana (Eaton) Clute, Fern Allies, 258 (1905); Isoetes engelmanni var. gracilis Engelm. in A.Gray, Manual of Botany of the Northern United States ed. 5, 677 (1876); Isoetes engelmannii var. valida Engelm. in A.Gray, A Manual of Botany of the Northern United States (ed. 5) 677 (1867); Isoetes lacustris var. microspora Engelm., A. Br., Flora 29: 179 (1846); Isoetes engelmanni var. georgiana Engelm., Transactions of the Academy of Science of St. Louis 4: 384 (1882); Isoetes engelmannii forma georgiana (Engelm.) Clute, Fern Allies 258 (1905) - Eastern Canada (Ontario), eastern USA (Alabama, Arkansas, Connecticut, Delaware, Florida, Georgia, Illinois, Indiana, Kentucky, Maryland, Massachusetts, Michigan, Missouri, New Hampshire, New Jersey, New York, North Carolina, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Vermont, Virginia, West Virginia)]. Map Muenscher: 343, Fl. N Amer.: 70. Ill. Taylor & Hickey, 1992: 618, AWP 1: 26, Muenscher: 341, AWP 1: 26, Crow & Hellquist 1: 26.

Isoetes engelmannii x I. riparia [Isoetes brittonii D.F. Brunton & W.C. Taylor, 1990, Amer. Fern J. 80 (3): 82-84 (I. engelmannii A.C. H. Braun)]

Isoetes erongensis Wanntorp, 1970, Svensk bot. Tidskr. 64 (2): 150-152 - Swaziland, Namibia.

Isoetes eshbaughii Hickey & Fuchs, 1986, System. Bot. 11 (2): 310 - Bolivia: Cordillera de Cochabamba, 4420 m. Ill. Hickey, 1986: 311, 313; ill.  microspores  MUSSELMAN, 2002.

Isoetes escondidensis Halloy, 1980, Lilloa, Rev. bot. 35 (2): 67-l75, ill. [I. filipovichii Fuchs, nom. nud.; I. castillonii Fuchs nom. nud., Isoetes filipovichii Fuchs-Eckert, 1965, nomen nud.] - Argentina: Catamarca, Tucumán: Laguna Escondida Media, 4330 m, Laguna Perdida, 4500 m, Laguna Muerta, 4260 m, Laguna de la Manza, 4200 m, Quebrada de los Cazadores, 4500 m. Ill. microspores MUSSELMANN, 2002.

Isoetes favulata Ill.  microspores  MUSSELMAN, 2002.

Isoetes feuereri Fuchs-Eckert, 1985, nom. nud. Locus class.: Bolivia: Camacho, 4100 m  (?= I. boliviensis)

Isoetes filiformis Fuchs-Eckert, 1979, nomen nud. Locus class.: Brasil: Rio Grande do Sul: São Leopoldo = I. gonzagensis = I. brasiliensis.

Isoetes filipovichii Fuchs-Eckert, 1965, nomen nud. Locus class.: Argentina: Catamarca: Quebrada de Los Cazadores, 4500 m = I. escondidensis

Isoetes flaccida (Shuttlew.) A. Br., Flora 29: 178 (1846) [Calamaria flaccida (Shuttlew.) Kuntze, Rev. Gen. Pl. 2. 828 (1891); Isoetes alata Small, Ferns of Florida 133 (1918) ?, apud Pfeiffer, Ann. Mo. Bot. Gard. 9: 137 (1922); Isoetes chapmanii (Engelm.) Small, Ferns of Florida, 222: (1931) 1932; Isoetes flaccida forma chapmanii (Engelm.) Clute, Fern Allies, 258 (1905); Isoetes flaccida forma rigida (Engelm.) Clute, Fern Allies, 258 (1905); Isoetes flaccida var. alata (Small) N. Pfeiffer, Annals of the Missouri Botanical Garden 9 (2): 137 (1922); Isoetes flaccida var. chapmanii Engelm., Transactions of the Academy of Science of St. Louis 4: 386 (1882); Isoetes flaccida var. rigida Engelm., Trans. St. Louis Acad. Sci. 4: 386 (1882)] - C Florida, N Florida, S Georgia. Map Muenscher: 343, Fl. N. Amer.: 69.

Isoetes fluitans M.I. Romero, 2004 Bot. J.  Linnean Soc. 146 (2): 231-236 (Isoetes longissima sensu Prada, 1983: 85).  2n = 44 - Spain: Galicia, rivers.

Isoetes fontana = I. engelmannii

Isoetes foveolata A.A. Eaton in Dodge. Isoetes foveolata var. plenospora A.A.Eaton = I. tuckermannii x I. engelmannii

Isoetes frigida Croft [ined.] [Isoetes stevensii auct. non Croft: Croft (1980)] - Only known from around a single small lake on the southern side of Mt Saruwaged. http://www.anbg.gov.au/fern/aquatic/isoetaceae.html

Isoetes fuliginosa R.L.Small & R.J. Hickey, 2001, Amer. Fern J. 91(2): 55-57, figs. 5-7, 14-15. 2n = 44 (Isoetes arumiana Fuchs, 1959, nom. nud. Proc. Kon. Ned. Adad. Wetensch. C. 85 (2): 259) - Colombia: Cauca: Cordillera Central: Lagunilla de Las Casitas, 3700 m; only known from type locality. [?= I. cusiyacoensis - Laguna de Cusiyaco, 3200 m].

Isoetes fusco-marginata H.P. Fuchs, 1986, Fl. Illustr. Catar. 1 (Isoetaceas): 12 (I. isabelloccensis Fuchs, 1982) - Brazil: Rio Grande do Sul: São Francisco de Paula, 900 m, known only from type locality. Ill. Fuchs, 1986: 14; MUSSELMAN, 2002 (microspores). ?= I. organensis

Isoetes gardneriana Kunze ex Mett., 1859, Fil. Lechl. Fasc. 11: 36) [Calamaria gardneriana (Kuntze ex Mett.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Calamaria martii (A. Br.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes martii A. Br. ex Kuhn) Martius, Fl. Bras. 1 (2): 646, t. 78 (1884); Calamaria amazonica (A. Braun acc. Kuhn in Martius) Kuntze, Rev. Gen Pl. 2: 828 (1891); I. amazonica [Baker, Journ. Bot. 18: 109 (1880)], I. amazonica auct. non A. Br.; I. ramboi auct., Isoetes ramboi forma latifolia Herter)] - Brazil: Minas Gerais, Rio Grande do Sul: Santa Catarina (I. martii); Goiàs, Maranhão: Ilha da Balsas; Parà: Santarém; Rio de Janeiro (Itatiaia: Campo do Silveria - Fuchs-Eckert, 1982: 246), Paraguay, Argentina: Capurro. Map Fuchs-Eckert, 1986: 20, ill.: 14. Ill.  microspores  MUSSELMAN, 2002.

Isoetes gardneriana sensu A.C. Brade = I. bradei

Isoetes garnieri = I. nigritiana

Isoetes gemmifera Rauh, 1959 = I. andicola

Isoetes georgiana Luebke, Amer. Fern J. 82: 14, 1992 = I. boomii

Isoetes giessii Launert, 1957, Mitt. bot. Staatssamml. München 2: 309 (I. perrieriana auct. non Iversen) - Swaziland; Namibia: Erongo Mts. Ill. COOK, 2004: 34.

Isoetes gigantea U. Weber, 1922, Nova Hedwigia 63: 245 - Rio de Janeiro, W Bahia. [Isoetes graminifolia Fuchs-Eckert, 1980, nomen nud. (=  I. vannielii, I. rosselii Fuchs, ined. - W Bahia). Locus class.: Brasil: Mato Grosso (in rapidly flowing water)] - Maranhão, Bahia: rio das Ondas, 4 km N Barreiras, 550 m (Fuchs-Eckert, 1982: 247), Lagoa Itaparica. Map Fuchs-Eckert, 1982].

Isoetes glacialis Asplund, 1926 - Columbia: Cundinamarca, Boyacá, Cauca, Meta; Bolivia: La Paz; Peru: Junín, Puno = I. lechlerii Mett.

Isoetes goebelii H. Christ, ined., in Fuchs-Eckert, 1992: 249 = I. ovata Pfeiffer

Isoetes goebelii Weber, 1922, Hedwigia 63: 247, fig. 16-17 - Brazil: Paraíba, Rio de Janeiro (in sectio Isoetes, Fuchs, 1982).

Isoetes gonzagensis Fuchs-Eckert, 1979, nomen nud. (I. weberi sensu Dutra in sched.) Locus class.: Brasil: Rio Grande do Sul: São Leopoldo = I. filiformis = I. brasiliensis.

Isoetes gracilis = I. engelmannii

Isoetes graminifolia Fuchs-Eckert, 1980, nomen nud. Locus class.: Brasil: Mato Grosso (in rapidly flowing water) - Maranhão, Bahia: rio das Ondas, 4 km N Barreiras, 550 m (Fuchs-Eckert, 1982: 247), Lagoa Itaparica. Map Fuchs-Eckert, 1982 (=  I. van-nielii, I. rosselii) ?= I. gigantea Weber, 1922.

Isoetes grammontii = I. velata

Isoetes gravesii A.A.Eaton = I. setacea subsp. braunii

Isoetes gunnii A. Braun, 1868, Monatsber. Königl. Preuss. Akad. Wiss. 535 [Calamaria gunnii (A. Br.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes lacustris (L.) Rodway, Tasm. Fl. 279 (1903)] - Tasmania. Map Marsden, 1979: 337.

Isoetes guyanae U. Weber, nom. nud. (Fuchs-Eckert, 1982). Locus class.: Brasil: Amazonas: Serra do Mel, Rio Branco = I. triangula

Isoetes gymnocarpa (Gennari) Braun = I. histrix

Isoetes habbemensis Alston, 1945, J. Arn. Arb. 26 - Irian Jaya: Mt. Wilhelmina: Lake Habbema, and Papua New Guinea: Star Mts., 3200-3300 m. Map Croft, 1980: 182 Ill. Croft, 1980: 178, 185.

Isoetes hagemannii Fuchs-Eckert, 1981, nomen nud. Locus class.: Colombia: Nariño: Páramo del Volcán Galeras, 4000-4200 m.  ?= I. novogranadensis

Isoetes halloyi Fuchs-Eckert, 1981, nomen nud. Locus class: Argentina: Tucumán: Laguna Perdida, 4500 m, Laguna Escondida Media, 4330 m. = I. escondidensis

Isoetes harveyi A.A.Eaton = I. lacustris x I. tuckermannii

Isoetes hawaiiensis W.C.Taylor & W.H.Wagner, 1993, Amer. Fern J. 83(2): 67 - Hawaii (Maui, Hawaii Isl.)

Isoetes heldreichii Wettst., 1886, Verhandl. der Kais.-König. Zool.-Bot. Wiss. Wien 36: 239, t. 8 - C Greece (Paleokastron), 1050 m. Ill. C & K 1: 52, 55. = prob. I.  delilei subsp.

Isoetes hemivelata R.L. Small & Hickey, 2001, Amer. Fern J. 91(2): 57 (60, figs. 9-11, 16-17). 2n = 44 - Colombia: Cordillera Central, cabeceras de los ríos Tulua y Bugalagrande: Páramo de Las Vegas, 3600-3800 m (type); dept. Huila/Cauca: Macizo Colombiano; páramo de Las Papas, Cerros y alrededores de la laguna La Magdalena, 3530 m; Meta: páramo de Sumapaz, Hoya Sitiales, Laguna La Primavera, 3425 m; Risaralda: Cordillera Central; Municipie de Pereira; parque de los Nevados, 4250 m; dpart. del Valle: páramo Pan de Azúcar, 3300-3700 m. Submerged in permanent lakes and ponds, occasionally streams 3300-4300 m.

Isoetes herteri Fuchs, 1960, nom. nud. Locus class.: Uruguay: Dept. de Rocha: Matavillas =? I. smithii

Isoetes herzogii Weber, 1922, Hedwigia 63: 250, fig. 22-23 - Perú: Cuzco; Bolivia: Cochabamba. Ill.  microspores  MUSSELMAN, 2002.

Isoetes heterospora A.A.Eaton, 1900, Fernwort Papers 8 = I. lacustris

Isoetes hewitsonii Hickey, 1994, Fieldiana, Bot. 34: 94 - Andes of Perú: Cajamarca, 3700 m, known only from type loc.

Isoetes hickeyi Fuchs-Eckert, 1981, nomen nud. Locus class.: Venezuela: Bolivar: Cima del Roraima.

Isoetes hickeyi Taylor & Luebke = I. setacea x I. lacustris

Isoetes hieroglyphica A.A.Eaton = I. lacustris

Isoetes hieronymi Weber, 1922, Hedwigia 63: 251, fig. 24-25 (I. argentina Eaton & Palmer, Amer. Fern Journ. 19: 18 (1929) nom. nud., I. kanteri auct. non R. Brown, I. lechleri Pfeiffer, 1922, non Mett., I. socia auct. non Br.) - Argentina: Córdoba; Chile: Nahuelbuta, Coquimbo, Norte Chico. Ill.  microspores  MUSSELMAN, 2002.

Isoetes histrix Bory, 1844, Compt. Rend. Hebd. Séances Acad. Sci. 18: 1167. 2n = 20 (2n = 22). [Isoetes histrix Bory var. histrix; Isoetes hystrix Wolsey, New Phytol. 11. 5: 45 (1861); Isoetes hystrix Gdgr., Fl. Eur. 27: 197 (1891); Calamaria hystrix (Dur.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Cephaloceraton hystrix Gennari, Comment. Critt. Ital. 1 (3). 111 (18 Comp. Fl. lt., 25 (1867), et Motel. & Vendr., Actes Soc. Linn. Bord. 36: 4009, t. 17, f. 1-5 (1883); 62); Isoetes chaetureti, Mendes, Agron. Lusit. 23: 7 (1961);  Isoetes delalandei Lloyd, Notes Fl. Ouest France: 25 (1851); Isoetes histrix subsp. delalandei (Lloyd) P. Fourn., Quatre Fl. France: 6 (1934); Isoëtes histrix proles delalandei (J. Lloyd) Rouy [1913, Fl. Fr. 14: 481; I. histrix subsp. delalandei (J. Lloyd) P. Fourn., 1934, Quatre Fl. Fr.: 6; Isoetes duriaei Hook., Brit. Ferns, t. 56 (1861); Isoetes histrix forma caespitosa Motel. & Vendr., Actes Soc. Linn. Bord. 36: 401 (1883); Isoetes histrix forma grandis Motel. & Vendr., Actes Soc. Linn. Bord. 36: 402 (1883); Isoetes histrix forma loricata (a) grandis A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec. (1863): 617 (1864); Isoetes histrix forma loricata (b) nana A. Br., Monatsber. K. Acad. Wiss. Berlin, Sitzung 7 Dec. (1863): 617 (1864); Isoetes histrix forma loricata A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec. (1863): 617 (1864); Isoetes histrix forma nana Motel. & Vendr., Actes Soc. L. Bord. 36: 402 (1883); Isoetes histrix forma solitaria Motel. & Vendr., Actes Soc. L. Bord. 36: 401 (1883); Isoetes histrix var. submersa Batt. & Trab., Fl. Alg. & Tunis., 407 (1902); Isoetes histrix forma submersa (Trabut) Maire, Jahandiez & Maire, Cat. Pl. Maroc. 1: 12 (1931); Isoetes histrix var. grandis Milde, Batt. & Trab., Fl. Alg. & Tunis. 407 (1902); Isoetes histrix var. lacunosa Tod., Syn. Pl. Acot. Vasc. Sic., p. 46 (1866); Isoetes histrix (1863), et Batt. & Trab., Fl. Alg. & Tunis., 407 (1902); Isoetes histrix var. loricata subvar. armata Batt. & Trab., Fl. Alg. et Tunis., 407 (1902); Isoetes histrix var. loricata subvar. inermis Batt. & Trab., Fl. Alg. et Tunis., 407 (1902); Isoetes histrix var. scutellata A. Br. apud A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec. (1863): 617 (1864), Arm. Sci. Nat. 2: 371 (1864); Isoetes histrix var. typica Fiori & Paoletti, Fl. Anal. Ital. 1: 25 (1896); Isoetes hystrix algarbiensis Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix arcuata Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix cesatii Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix collina Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix corsica Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix horridula Gdgr. Fl. Eur. 27: 197 (1891); Isoetes hystrix Iongifolia Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix longiseta Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix numidica Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix oceanica Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix phrygia Gdr., Fl. Eur. 27: 197 (1891); Isoetes phrygia Hausskn., Mitth. Thüring. Bot. Vereins 13: 72 (1899); Isoetes hystrix todaroi Gdgr., Fl. Eur. 27: 197 (1891); Isoetes hystrix transiens Gdgr., Fl. Eur. 27: 197 (1891); Isoetes setacea Moris & DeNotaris, Florula Capr. 161 (1839) - Atlantic Europe: England, France, Portugal; Spain, Corsica, Sardinia, Malta, Sicily, Italy, Tunisia, Algeria, Morocco, former Yugoslavia, central Macedonia, Greece, Crete, Aegaean Isl., Turkey (Anatolia). Ill. C & K 1: 52, 59, Prelli: 83, BERTHET & LECOCQ, 1977: 331, 341, 342; BOLIN et al. 2008. Cephaloceraton pseudohystrix Genn., Ann. Agric. Sic. An. XI): 175 (1865); Isoetes pseudohystrix Genn., Levier, Barb., Fl. Sardoae Comp. 74, 195 (1885); - N Mediterranean east to Turkey, S Syria and N Lebanon. Ill. BOLIN et al., 2008.

Isoetes hookeri A.Braun = I. humilior

Isoetes hopei J.R. Croft, 1980, Blumea 26: 177-190, map: 182 - Irian Jaya: Mt. Carstenz.. Map Croft, 1980: 179-185 (terrestrial).

Isoetes howellii Engelm., 1882, Trans. Acad. Sci. St. Louis 4: 385. 2n = 22 [Isoetes howellii var. piperi Clute; Isoetes melanopoda var. californica A.A. Eaton in Gilbert; Isoetes melanopoda var. californica A.A. Eaton, Working List of North American Pteridophytes 27 (1901); Isoetes nuda Engelm., Trans. St. Louis Acad. Sci. 4: 385 (1882); Isoetes underwoodii L.F. Henderson, Botanical Gazette 23: 124 (1897); Isoetes minima Eaton] - S British Columbia, W Montana, Idaho, Washington, Oregon, California to Mexico (Baja California). Map Fl. N Amer.: 74, ill.: 71, map Muenscher: 343. Ill. Taylor & Hickey, 1992: 615.

Isoetes humilior F. Muell. ex A. Br., 1852, Linnaea 25: 722 [I. tasmanica Durieu, 1864; Calamaria humilior (F. Muell. ex A. Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes hookeri A. Braun (in Herb. Melbourne (1866), Monatsber. K. Akad. Wiss. Berlin, 538 (1868)] - Central Tasmania. Map Marsden, 1979: 338.

Isoetes hyemalis D.F.Brunton & Britton, 1994, Castanea 59(1): 13 - North Carolina, South Carolina, Virginia, Florida = I. saccharata = I. riparia

Isoetes hypsophila Handel-Mazzetti, 1923, Anzeiger Acad. Wiss. Wien. Nr. 13: 95 (2n = 22) - China: Yunnan.

Isoetes hystrix Durieu = I. histrix Bory

Isoetes hystrix hispanica Gdgr., Fl. Eur. 27: 197 (1891) = I. histrix

Isoetes hystrix subinermis Gdgr., Fl. Eur. 27: 197 (1891) see I. histrix

Isoetes hystrix oranensis Gdgr., Fl. Eur. 27: 197 (1891) = I. histrix

Isoetes immaculata Maire, 1952, pro forma I. velatae, subsp. intermediae = I. velata

Isoetes incerta Hollick, 1928 - Puerto Rico (Tertiary).

Isoetes indica Pant & Srivastava, 1962, The genus Isoetes in India. Proc. Nat. Inst. Sci. India, vol. 28, part B: 242-280 - India: Madhya Pradesh, known from type locality only. Ill. loc. cit. 269-272, map: 277 = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes inflata E.R.L. Johnson, 1984, J. Roy. Soc. W. Australia 67(1): 37 - SW Western Australia. Map Marsden, 1979: 335.

Isoetes intermedia (Trabut, pro var. I. velatae) Maire & Weiller, 1952 = I. velata

Isoetes isabellocensis Fuchs, 1960, nomen nud. Locus class.: Brasil: Rio de Janeiro: Serra dos Orgãos: Isabelloca, 2000 m. = I. itatiaiae, = I. braunii = I. organensis

Isoetes itaboensis Fuchs-Eckert, 1981, nomen nud. Locus class: Paraguay: Río Itabó, near mouth of Río Paraná. Ill.  microspores  MUSSELMAN, 2002. (= I. paranaensis ?= I. smithii)

Isoetes itatiaiae G. Herter, nom. nud., pro syn. I. dusenii. Locus class.: Brasil: Serra do Itatiaia, 2100 m. = I. braunii = I. isabelloccensis = I. organensis

Isoetes jaegeri Pitot, 1968, Bull. Inst. Fond. Afr. Noire, A, 30: 58, f. 1-19 - Burkina Faso: Haute Volta: plateau de Touziana.

Isoetes jamaicensis Hickey, 1981, Amer. Fern J. 71(3): 69. 2n = 22 - Jamaica: Harris Savanna.

Isoetes japonica A. Br., 1861, Monatsber. K. Akad. Wiss. Berl. 1: 459 [Calamaria japonica (A. Br. ) Kuntze, Rev. Gen. Pl. 2: 828 (1891)]; Isoetes edulis Sieb. ex Miq., Prol. Fl. jap. 390 (1866)-67 - Japan, Yunnan, Korea. I. j. subsp. japonica - Japan. I. j. subsp. sinensis Palmer [Isoetes sinensis var. coreana (Y. H. Chung & H. K. Choi) M. Takamiya & al.] - China: Kiangsu, near Nanking, Korea and S Japan. Maps MARSDEN, 1979: 342, map and ill. Kadono: 8, 10, ill. Chinese aq.: 19; UEHARA, 1991.

Isoetes jeffreyi = I. lacustris x I. riparia

Isoetes jsidis U. Weber, nomen nud. Locus class.: Brasil: Rio Grande do Sul: Porto Alegre, 100 a.s.l.  = I. dusenii

Isoetes junciformis Brunton D.F. & Britton D.M., 1999, Amer. Fern J. 89 (3): 187-197 - S Georgia.

Isoetes karstenii A. Braun, 1862, Verh. Bot. Vereins Prov. Brandenburg 3-4: 332. Type: Venezuela: Mérida, 2400 m. 2n = 22 (?= I. pichichensis). Isoetes karstenii var. karstenii [I. steyermarkii Fuchs, 1960, nom. nud., Proc. Koninkl. Ned. Acad. Wetensch. C 85: 257, 1982; I. colombiana (Palmer) Fuchs-Eckert, 1981, Proc. Kon. Nederl. Akad. Wetensch. C 84(2): 176, Isoetes lechleri var. colombiana T.C. Palmer, Amer. Fern J. 19: 18, 1929] - Locus class.: Venezuela: Mérida, 2400 m, submerged in permanent lake; Mérida, near El Aquila, 3930 m. Also: Mérida, 3400, 4000 m; Colombia: Santander: Laguna de Cunta, edge of Páramo de Santurbán, 3880 m; Meta, Tolima, Nariño; Boyacá, Arauca, Cundinamarca, Cauca, Huila-Cauca, dept. del Valle, 3530-4000 m. Isoetes karstenii var. anomala (T.C. Palmer) R.L. Small & Hickey American Fern Journal 91 (2): 48 (2000) [Isoetes rimbachiana Fuchs-Eckert, Proc. Kon. Nederl. Akad. Wetensch. C 85(2): 226 1982, Proc. Kon. Nederl. Akad. Wetensch. 85 (2): 226] - Known only from the type locality Mount Chimborazo, shallow pond 4200 m, although Rodríguez (1955) cites one specimen of I. lechleri var. anomala from Laguna Negra, Mérida, Venezuela (Small & Hickey). (sectio Laeves - Fuchs, 1982). 

Isoetes keeleyorum Fuchs-Eckert, 1988, nomen nud. Locus classs: Ecuador: Napo, North of Quito, 3850 m (? = I. ecuadoriensis)

Isoetes kersii Wanntorp, 1970, Svensk bot. Tidskr. 64 (2): 146 - S Africa: Swaziland, Natal.

Isoetes killipii C.V. Morton, 1945, American Fern Journal 35: 49 (I. bischlerae Fuchs in sched. - Boyacá, Cundinamarca, Antioquia, Cauca; I. hagemanni Fuchs in sched. - Nariño; I. roraimensis Fuchs, nom. nud.) - Colombia, 2100-4000 m (Páramo de Puracé, Boyacá, Valle, Cauca, Nariño, Charalà, Pàramo de la Rusia), Venezuela, Ecuador;  British Guyana: Mt. Roraima (I. roraimensis), Brazil: Serra Pacaraima 2740 m.. 

Isoetes kirkii A. Br., 1869, Monatsber. K. Akad. Wiss. Berlin. 649. 2n = 22 [Calamaria kirkii (A. Br.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)] - New Zealand (North Isl., South Isl.)] I. k. var kirkii, I. k. var flabellata. Map Marsden, 1979: 339. Ill. Johnson & Brooke: 31, Aqua-Planta 2000 (4): 136. However  see I.  alpina.

Isoetes kriegeri Fuchs-Eckert, 1980, nom. nud. Locus class: Brasil: Minas Gerais: Parque Nacional do Caparaó, 2400 m ?= I. smithii

Isoetes lacustris (L.) Rodway = I. gunnii

Isoetes lacustris circinnata, curvifolia, echinospora, longifolia, minor, patentissima, patula, rectifolia, recurva, recurvata, terrestris, stricta = I. setacea Lam.

Isoetes lacustris Engelm. = I. lacustris L.

Isoetes lacustris L. sensu Bert. = I. setacea Lam.

Isoetes lacustris L. sensu Moris = I. velata

Isoetes lacustris L., 1753, Sp. Pl.: 1100. 2n = c. 110 [Calamaria lacustris (L.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); I. macrospora Durieu; Isoetes atrovirens Th. Fr., Bot. Not. (1863): 48 (1865); Isoetes harveyi Eaton, Fernwort Papers 2: 11 (1900); Isoetes hieroglyphica A.A. Eaton, Fernwort Papers 10 (1900); Isoetes macrospora forma hieroglyphica (Eaton) N.E. Pfeiffer; Isoetes macrospora forma hieroglyphica (A.A. Eaton) N.E. Pfeiffer, Annals of the Missouri Botanical Garden 9(2): 194 (1922) Isoetes lacustris subsp. hieroglyphica (A.A. Eaton) Taylor & Luebke; Isoetes lacustris abreviata Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris arvernensis Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris cambrica Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris editorum Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris Engelm., Trans. St. Louis Acad. Sci. 4: 377 (1882); Isoetes lacustris errabunda Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris forma curvifolia subf. falcata Tausch, Schmidt, Unterrichtsanst. Klosters St. Johannis zu Hamburg 1902-03: 74 (1903); I. lacustris subsp. echinospora (Durieu) Moore & More, 1866, Contr. Cybele Hibern.: 381; Isoetes lacustris forma subfalcata Gay, Bull. Soc. Bot. France 10: 422 (1863); Isoetes lacustris forma semifalcata Gay, Bull. Soc. Bot. France 10: 423 (1863); I. lacustris var. falcatum Fliche, 1879, Les Isoètes des Vosges: 7; Isoetes lacustris forma elatior Fliche, Les Isoètes des Vosges, Mem. Acad. Stanislas. IV. 11 (1878): 181 (182) (1879); Isoetes lacustris forma echinospora b. elatius Fliche, ex Fiori & Paoletti, App. Fl. Anal. Ital. 10 (1907-1908); I. lacustris var. minor A. Br. ex Milde,1865, Sporenfl.: 141; I. lacustris var. strictum J. Gay;1863, Bull. Soc. Bot. Fr., 10: 392; Isoetes lacustris forma gemmifera Mer, Bull. Soc. Bot. France 28: 72 (1881); Isoetes lacustris forma maxima Alex. Blytt, Norges Flora 111 (Suppl.): 1238, 1876; Isoetes lacustris var. paupercula Engelm.; Isoetes lacustris subsp. paupercula (Engelm.) Feilberg. Trans. St. Louis Acad. 4: 377 (1882); Isoetes lacustris forma stricta Gay, Bull. Soc. Bot. France 10: 392, 424 (1863); Isoetes lacustris forma tenuifolia A. Br., Milde, Fil. Eur. & Atl. 277 (1867); Isoetes lacustris forma terrestris Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 11, fig. 5 (1936); Isoetes lacustris forma typica Fiori & Paoletti, Fl. Anal. Ital. 1: 25 (1896); Isoetes lacustris lamottei Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris langei Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris latifolia Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris noltei Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris obscura Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris pervaga Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris Pursh, Fl. Amer. Sept. 671 (1814); Isoetes lacustris rivalis Gdgr., Fl. Eur. 27. 193 (1891); Isoetes lacustris rossica Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris scanica Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris scotica Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris septentrionalis Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris spreta Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris subsp. lacustris Hook. forma, Student's Fl. Brit. Isl., ed. 1, 471 (1870); Isoetes lacustris succedanea Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris suecica Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris var. curvifolia Glück, Die Süsswasser-Flora Mitteleuropas 15: 10 (pro syn.) (1936); Isoetes lacustris var. humilior Motel. & Vendr., Actes Soc. Linn Bord. 36: 327 (1883); Isoetes lacustris var. humilis Mer, Bull. Soc. Bot. Fr. 28- 72 (1881); Isoetes lacustris var. intermedia Mer, Bull. Soc. Bot. Fr. 28: 72 (1881); Isoetes lacustris var. leiospora Klinggraff., Schrift. Nat. Ges. Danzig, Vl. 1: 20 (1884); Isoetes lacustris var. morei (Moore) Hooker, J. D., Student's Fl. Brit. Isl., ed. 3, 526 (1884); Isoetes lacustris var. rectifolia Caspary, Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 10 (1936); Isoetes lacustris var. recurva Motel. & Vendr., Actes Soc. L. Bord. 36: 327 (1883); Isoetes lacustris var. setacea Leman, Dict. Sci. Nat. 24: 24 (1822); Isoetes lacustris var. stricta (Gay) Mer, Bull. Soc. Bot. Fr. 28: 72 (1881); Isoetes lacustris var. tenuifolia A. Br. ex Milde, Sporenpfl. 141 (1865); Isoetes lacustris var. terrestris Motel. & Vendr., Actes Soc. Linn. Bord. 36: 327 (1883); Isoetes lacustris var. vulgaris Casp., Luerss., Farnpfl., 855 (1889); Isoetes lacustris vergens Gdgr., Fl. Eur. 27: 193 (1891); Isoetes lacustris x stricta Gay, Bull. Soc. Bot. France 10: 392, 424 (1863); Isoetes leiospora Klinggraff., Schrift. Nat. Ges. Danzig 6 (1): 20 (1884); Isoetes macrospora Dur., Bull. Soc. Bot. Fr. 11: 101 (1864); Isoetes macrospora forma heterospora (Eaton) Clute, Fern Bull. 16: 56 (1908); Isoetes heterospora Eaton, Fernwort Papers, 2: 8 (1900); Isoetes macrospora var. heterospora Eaton, Rhodora 10: 42 (1908); Isoetes morei D. Moore, Journ. Bot. Lond. 16 (n. s. 7): 353, t. 199 (1878); Isoetes tuckermani var. harveyi (Eaton) Clute, Fern Allies, 226, 227, 260 (1905); Isoetes tuckermani var. heterospora (Eaton) Clute, Fern Allies, 226, 260 (1905)] - Iceland, Norway, Sweden, Finland, N and C European Russia, Baltic States, Ireland, England,  Denmark, Netherlands, Belgium, France, N Spain, Germany, Switzerland, Poland, Austria, former Czechoslovakia, ?Romania, (disjunct) Bulgaria (Pirin Mts., W Rhodopes), Greece, W Siberia, Altai, C Siberia (and, disjunct, Japan ?), S Greenland, C and E Canada (Manitoba, New Brunswick, Labrador, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Quebec, Saskatchewan), northeastern USA (Connecticut, Maine, Massachusetts, Michigan, Minnesota, New Hampshire, New York, Rhode Island, Tennessee, Vermont, Virginia, Wisconsin). Map Hultén & Fries 1: 5, Fl. N Amer.: 68, ill. C & K 1: 52, Prelli: 83, Fassett: 46, AWP 1: 26, Crow & Hellquist 1: 26; BERTHET & LECOCQ, 1977: 335, 353, 354.

Isoetes lacustris Pursh. = I. lacustris L.

Isoetes lacustris sensu Hook., 1861, Brit. Ferns = I. amazonica (part), I. gardneriana (part)

Isoetes lacustris var. microspora = I. engelmanni

Isoetes lacustris x I. riparia (I. jeffreyi)

Isoetes lacustris x I. tuckermannii (I. x harveyi)

Isoetes laevis U.Weber = I. lechleri

Isoetes lancifolia Fuchs-Eckert, 1981, nom. nud. Locus class.: Ecuador: Napo: Laguna.

Isoetes lechleri (Mettenius) A. Braun, 1862 [Calamaria lechleri Mettenius, Fil. Lechler. 2: 36, 1859; Calamaria lechleri (Mettenius) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes socia A.Braun, 1862, Verh. Bot. Ver. Brandenbg. 3-4: 332; Isoetes glacialis Asplund, 1926; Isoetes laevis U.Weber Hedwigia 63: 252, fig. 30-31 (1922); Isoetes peruviana Weber, Hedwigia 63: 246, fig. 12-13 (1922); Isoetes socia A. Br., Verh. Bot. Ver. Brandenb. 4: 332 (1862); Isoetes lechleri var. triquetra (A. Br.) L.D. Gómez, Brenesia 18: 5, 1981] - Andes: Venezuela (Mérida: Sierra Nevada de Santo Domingo; Distrito Federal: Pico de Naiguatá), Colombia (Santander, Huila, Cauca, Arauca, Boyacá, Meta, Cundinamarca), Ecuador (Chimborazo, Pinchincha, Napo), Bolivia (La Paz) to Perú (Junín, Puno: Ayapata; Ayacucho, Ancashi: Cordillera Negra, 4400 m above Caras) 2400-4750 m. Ill. Tryon, 1982: 83; MUSSELMAN, 2002 (microspores).

Isoetes lechleri Pfeiffer, 1922, Ann. Missouri Bot. Garden 9 (2): 138 = I. hieronymi Weber

Isoetes lechleri var. colombiana T.C. Palmer, Amer. Fern J. 19: 18, 1929 = I. karsteni var. karsteni

Isoetes lechleri var. savatieri (Franch.) L.D. Gómez = I. savatieri

Isoetes lechleri var. triquetra (A.Br.) L.D. Gómez = I. triquetra

Isoetes leiospora Klingraff = I. lacustris

Isoetes leistii Fuchs-Eckert, 1981, nomen nud. Locus class.: Colombia: Nariño: Cumbal: Laguna de Bolsa, 3400 m (? = I. novogranadensis)

Isoetes lereschii Reich., 1881 (pro var. I. boryanae) = I. velata subsp. velata

Isoetes lineolata = I. velata adspersa

Isoetes lithophila Pfeiffer, 1922, Ann. Missouri Bot. Gard. 9: 135 (I. lithophylla) - SC Texas: Edwards Plateau (Burnett Co., Mason Co., Llano Co.). Ill. Fl. N Amer.: 71, SWUS 1: 38.

Isoetes longissima Bory de Saint-Vincent & Durieu de Maisonneuve, 1844, Comptes-r. hebd. séances Acad. Sc. Paris 18: (1165). Locus classicus: dans une des mares des forêts qui environnent le lac Houbéria au canton de La Calle (= El Kala, Algeria) = I. velata subsp. longissima

Isoetes longissima sensu Prada, 1983: 85 = Isoetes fluitans

Isoetes louisianensis John W. Thieret, 1973, Sida 5: 129, Sida, Contrib. Bot. 5: 129. SE Louisiana: Washington Parish. - SE USA. (I. louisianensis may represent I. engelmannii x melanopoda - Fl. N Amer.: 71).

Isoetes luetzelburgii Weber ex Luetzelburg, Estud. Bot. Nordeste 3: 257, 1926.  (Isoetes borboremensis Fuchs, 1960, nom. nudum) - Brazil: Rio Grande do Norte; Parahiba do Norte, Serra do Borborema: Lagoa dos Patos. Map Fuchs-Eckert, 1982.

Isoetes macounii A.A.Eaton = I. maritima

Isoetes macrospora Durieu, 1864 = I. lacustris L.

Isoetes macrospora forma hieroglyphica = I. lacustris

Isoetes macrospora forma paupercula = I. occidentalis

Isoetes mahadevensis Sristava, Pant & Shukla, 1993, Amer. Fern J. 83(4): 106 - India: Madhya Pradesh = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes malinverniana Cesati & De Not., Index Sem. Hort. Genuensis 1858: 36. 2n = 44 [Calamaria malinverniana (Ces. & De Not.) Kuntze, Revisio Generum Plantarum 2: 828 (1891) 1893; Isoetes malinverniana forma terrestris Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 14, fig. 7 (1936)] - NW Italy, very localized: Vercelli, Novara (streams in rice fields), Torino, streams (here apparently extinct). Map Cook, 1983: 560. Ill. C & K 1: 52, 55, Datz 48 (8): 526-529. Cf. Schneller, J.J., 1982.

Isoetes maritima Underw., 1888, Bot. Gaz. 13: 94 [Isoetes beringensis Komarov, Bull. Jard. Bot. Acad. Sci. U. S. S. R. 30 (1-2): 196 (1932), Isoetes echinospora subsp. maritima (Underw.) A.Löve; Isoetes muricata subsp. maritima (Underw.) Hultén. I. setacea subsp. maritima (Underw.) Holub, 1964, Acta Horti bot. prag.: 55; Isoetes echinospora subsp. maritima (Underw.) A. Löve; Isoetes muricata subsp. maritima (Underw.) Hult.; Isoetes echinospora forma maritima (Underw.) Clute; Isoetes echinospora var. maritima (Underw.) A.A.Eaton, Fern Bull. 13: 52 (1905); Isoetes braunii var. maritima (Underw.) Pfeiffer, Ann. Missouri. Bot. Gard. 9: 174 (1922); Isoetes echinospora var. braunii forma maritima (Underw.) Clute, Fern Bull. 16: 55 (1908); Isoetes echinospora var. hesperia (Reed) A.Löve; Isoetes echinospora var. savilei Boivin; Isoetes macounii Eaton, Fern Bull. 8: 12 (1900)] - Commander Isl., Aleutes, Alaska, W Canada (Vancouver Isl.), NW USA (Washington).

Isoetes martii A. Br., ex Kuhn in Martius, 1884, Fl. Bras. 1 (2): 646, t. 78 = I. smithii

Isoetes martii sensu Ule = I. triangula

Isoetes mattaponica Musselman & W.C.Taylor, 2001, Novon 11(2): 200-204, figs. 1-6)] - Mattaponica River, plants in Pontederia-Nuphar marsh, along a tidal shore of the Mattaponi River, ca. 0.75 km S of Aylett.

Isoetes maxima Sehnem, nom. nud. Locus class. Brasil: Rio Grande do Sul, Cambará: Fortaleza, 1200 m, in streams. Cf. I. brasiliensis

Isoetes melanopoda Gay & Dur., 1864, Bull. Soc. Bot. France 11: 102 [Calamaria melanopoda (Gay & Dur.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes virginica Pfeiffer, Claytonia 3: 29 (1937) - N Georgia, W South Carolina, C Virginia, northeastern Alabama (in Fl. N Amer., with piedmontana as syn.), Isoetes virginica var. piedmontana N. Pfeiff.; Isoetes melanopoda forma californica (Eaton) Clute, Fern Allies, 260 (1905); I. m. forma pallida (Engelm.) Fernald; Isoetes melanopoda var. pallida Engelm. Trans. St. Louis Acad. Sci. 4: 387 (1882)] - C USA: Arkansas, Georgia, Illinois, Indiana, Iowa, Kansas, Kentucky, Louisiana, Minnesota, Mississippi, Missouri, Nebraska, New Jersey, North Carolina, Oklahoma, South Carolina, South Dakota, Tennessee, Texas, Utah, Virginia. [Isoetes virginica Pfeiffer, Claytonia 3: 29 (1937) N Georgia, W South Carolina, C Virginia, NE Alabama, Ill. Fl. N Amer.: 71, SWUS 1: 38, AWP 1: 26, Crow & Hellquist 1: 26].

Isoetes melanopoda var. californica A.A.Eaton in Gilbert = I. howellii

Isoetes melanospora Engelm., 1877, Trans. Acad. Sci. St. Louis 3: 395 = Calamaria melanospora (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891). - C Georgia, South Carolina (Lancaster Co.). Ill. Fl. N Amer.: 71.

Isoetes melanotheca Alston, 1956, Bol. Soc. Brot., sér. 2A, 30: 15 (I. "melanothes ") - Guinea.

Isoetes mexicana Underw., 1888, Botanical Gazette 13: 93 [Isoetes montezumae A. A. Eaton, Fern Bulletin 5(2): 25 (1897)] - N Guatemala, Mexico, between 2200 and 2900 m (Chihuahua, Hidalgo, Morelos, Chiapas, Nayarit, Zacatecas, Aguascalientes, ?Guanajuato, Jalisco, Mexico State, Durango).

Isoetes meyeri Fuchs-Eckert, 1965, nom. nud. Locus class.: Argentina: Río Negro: Bariloche: Cascada del Río Manzo = I. savatieri

Isoetes microspora = I. engelmanni

Isoetes microvela D.F. Brunton & D.M. Britton, 1998, Rhodora 100 (903): 270-275, f. 1-3, 4 e-h, 5 - Coastal SE USA.

Isoetes minima A.A.Eaton, 1898, Fern Bulletin 6: 30. 2n = 22 [Isoetes howellii var. minima (Eaton) N. Pfeiff., Annals of the Missouri Botanical Garden 9(2): 142 (1922)] - Washington state, British Columbia. [Valid species, see Botanical Electronic News No. 269, June 2, 2001, and W.C. Taylor, N.T. Luebke & A.R. Lekschas, Taxonomic status and evolutionary relationship of Isoetes minima A.A. Eaton (Isoetaceae) based on nuclear ribosomal DNA internal transcribed spacer sequences, ibid. No. 304, March 6, 2003].

Isoetes minuata J. Drummond, 1846, in litt. ad W.J. Hooker (type lost). Locus class.: Western Australia: vicinity of Wangan.

Isoetes mirzapurensis Panigrahi & Dixit, 1965, Proc. Indian Sci. Congr. pt. III, Abstract, Botany 243 - India: Uttar Pradesh: Mirzapur district = I. indica Pant & Srivastava non J.G. König ex J.E. Smith = I. unilocularis = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes mongerensis E.R.L. Johnson, 1984, J. Roy. Soc. SW. Australia 67(1): 39 - SW Western Australia. Map Marsden, 1979: 336.

Isoetes montana T.C. Palmer, 1931 = I. storkii

Isoetes montana Weber, 1922, Hedwigia 63: 254, fig. 38-39 "Andes" (non Palmer, 1931). Nearest I. amazonica.

Isoetes montezumae A.A.Eaton = I. mexicana

Isoëtes morei T. Moore,1878, J. Bot. (London), 16: 353 = I. lacustris

Isoetes muelleri A. Br., 1868, Monatsber. K. Akad. Wiss. Berlin 541 [Calamaria muelleri (A. Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes stuartii A. Br., Monatsber. K. Akad. Wiss. Berlin, 539 (1868); Isoetes tenuissima F. Muell., 1868 (non Boreau) - All Australia and Tasmania. Map Marsden, 1979: 331.

Isoetes multiangularis = I. kirkii

Isoetes muricata Durieu = I. setacea Lam.

Isoetes muricata subsp. asiatica = I. setacea subsp. asiatica

Isoetes muricata subsp. truncata = I. maritima

Isoetes nana (Herter) Fuchs-Eckert, 1981 - Brasil: Serra do Itatiaia: Agulhas Negras, 2600 m = I. dusenii, ?= I. organensis

Isoetes narinensis Fuchs-Eckert, 1981, nom. nud. Locus class.: Colombia: Nariño: Cumbal: Laguna de Bolsa, 3400 m, also Laguna de Cumbal, 3500 m; Ecuador: Pichincha: Páramo de Guamani, Lojo, 2865 m, Napo-Pastaza, Cordillera de Los Llanganati, 3700 m = ? I. novogranadensis

Isoetes natalensis Baker, 1887 = I. welwitschii

Isoetes neoguineensis Baker ex F. v. M., 1898, Ann. Rep. Brit. N. Guinea 1897-8 (1898) [Isoetes neoguineensis Baker, Sadebeck, Nat. Pfl.-fam. 1 (4): 776 (1902); I. n. var. rheophila J.R. Croft, Blumea 26 (1980) 177-190, ill. 189] "Subsequent collections (Croft in press) have revealed intermediates between the two extremes so the variety can no longer be maintained, a decision foreshadowed by Marsden (1979)" - Papua New Guinea: Owen Stanley Ranges, 2800-4000 m (twice found). Map Croft, 1980: 182, ill.: 187.

Isoetes nigritiana A.Br. ex Kuhn, 1868 (I. garnieri Chevalier & Monnier, 1946) - Sudan: Bamako, Mali, Nigeria, Centrafrique, Ivory Coast. Ill.: Bull. Inst. Franç. Afrique noire, tome 16: 318, 1954.

Isoetes novogranadensis H.P. Fuchs, 1969, Caldasia 8: 314 [Isoetes dichotoma L.E. Mora & W. Hagemann, Mutisia 43: 5 (1977); Isoetes cf. killipii sensu Hickey apud Ollgaard A& Balslev, 1979, Rpts. bot. Inst. Univ. Aarhus (4): 431, non Morton, 1945, Amer. Fern J. 35 (2): 49; Isoetes cf. ramboi sensu Hickey apud Ollgaard & Balslev, 1979, Rpts. bot. Inst. Univ. Aarhus (4): 121; I. pichichensis, I. carchiensis and I. quilindanensis Fuchs in sched.] - Venezuela: (Mérida, Bolivar), Colombia (Valle, Huila, Boyacá, Cauca, Cundinamarca: de Villapinzón à Umbita; Putumayo, Páramo de San Antonio del Bordoncillo, 3250 m, Ecuador: Carchi, Pinchincha, Napo, Cotopaxi, Azuay; south  to Peru 3500-3600 m. [?= Isoetes narinensis Fuchs-Eckert, 1981, nom. nud. Locus class.: Colombia: Nariño: Cumbal: Laguna de Bolsa, 3400 m, also Laguna de Cumbal, 3500 m; Ecuador: Pichincha: Páramo de Guamani, Lojo, 2865 m, Napo-Pastaza, Cordillera de Los Llanganati, 3700 m; ?= Isoetes leistii Fuchs-Eckert, 1981, nomen nud. Locus class.: Colombia: Nariño: Cumbal: Laguna de Bolsa, 3400 m].

Isoetes nuda = I. howellii

Isoetes nuttallii A. Braun ex Engelm.,1874, Amer. Naturalist 8: 215. 2n = 22 [Calamaria nuttallii (A. Braun in Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Calamaria suksdorfii (Baker) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes opaca Nuttall, ex Engelm., Trans. St. Louis Acad. i. 4: 388 (1882); Isoetes suksdorfii Baker, Handbook of the Fern-Allies 132 (1887); - W USA (California, Oregon, Washington), W Canada (Vancouver Isl.). Map Fl. N Amer.: 74, ill.: 71, Taylor & Hickey, 1992: 615.

Isoetes occidentalis Henderson, 1900, Bull. Torr. Bot. Cl. 27: 358 (1900) [Isoetes echinospora var. braunii forma flettii (Eaton) Clute, Fern Bull. 16: 55 (1908); Isoetes echinospora var. flettii A.A. Eaton, Fern Bulletin 13: 51 (1905); Isoetes flettii (A.A. Eaton) N.E. Pfeiffer, Annals of the Missouri Botanical Garden 9(2): 186 (1922); Isoetes maritima var. flettii (A.A.Eaton) C.F.Reed; Isoetes echinospora forma flettii (A.A.Eaton) Clute; Isoetes echinospora var. flettii A. A. Eaton, Isoetes maritima var. flettii (Eaton) Reed, Amer. Fern Journ. 35: 85 (1945); (I. flettii has been attributed to both maritima and occidentalis); Isoetes piperi Eaton, Fern Bull. 11: 85 (nom. nud.) (1903); Isoetes occidentalis var. piperi (A.A. Eaton) A. Nelson & J.F. Macbr., Bot. Gaz. 61(1): 30 (1916); Isoetes howellii forma piperi (Eaton) Clute, Fern Bull. 16: 56 (1908); Isoetes howellii var. piperi (Eaton) Clute, Fern Allies, 246, 258 (1905); Isoetes occidentalis var. piperi A.Nelson & J.F. Macbr.); Isoetes lacustris var. paupercula Engelm., Trans. St. Louis Acad. Sc. 4: 377 (1882); Isoetes macrospora forma paupercula (Engelm.) Clute, Fern Bull. 16: 56 (1908); Isoetes occidentalis subsp. paupercula (Engelm.) Feilberg, Nordic J. Bot. 4(2): 155 (1984); Isoetes paupercula (Engelm.) A.A. Eaton, Proceedings of the United States National Museum 23(1226): 649 (1901); Isoetes lacustris subsp. paupercula (Engelm.) Feilberg. Trans. St. Louis Acad. Sc. 4: 377 (1882)] - Aleutes, Alaska, W Canada (British Columbia), NW USA (Washington, Oregon, C California, Idaho, Montana, Wyoming, Colorado). Map. Fl. N Amer.: 73, ill.: 71. (See I. lacustris and I. maritima)

Isoetes olympica A. Braun, Milde, 1867, Pl. Eur.: 285. 2n = 22 - Asiatic Turkey (Anatolia), S Syria: Hauran region. Remarks: I. histrix agg. = Calamaria olympica (A. Braun in Milde) Kuntze, Rev. Gen. Pl. 2: 828 (1891).  Ill. BOLIN et al., 2008. May be conspecific with I. histrix var. subinermis.

Isoetes opaca = I. nuttallii

Isoetes orcuttii Eaton, 1900, Fern Bull. 6: 30 (nom. nud.), 8: 13 (1900). 2n = 22 [Isoetes nuttallii var. orcutti (A.A. Eaton) Clute, The Fern Allies 253 (1905)] - C California to Baja California. Map Fl. N Amer.: 75, ill.: 71.

Isoetes organensis Weber apud Luetzelburg, Estud. Bot. Nordeste 3: 256, 1926 (I. braunii Weber, I. isabelloccensis, I. itatiaiae, I. dusenii) - Brazil: Rio de Janeiro, Serra dos Orgãos: Morro Açu, 2238 m, Isabellocca, 2000 m (in Fuchs-Eckert, 1982: 246).

Isoetes orientalis Liu Hong; Qing-Feng Wang & W. Carl Taylor, 2005, Novon 15 (1): 164-167. (2n = 66) - SE China: Zhejiang prov.: Songyang county, 1200 m.

Isoetes ovalifolia Fuchs-Eckert & Hickey, 1981, nom. nud. Locus class: Perú: Junín Cerro de Pasco, 4300 m. (= see I. asplundii)

Isoetes ovata Pfeiffer, 1922, Ann. Missouri Bot. Garden 8 (2): 109; loc. class.: British Guiana: pools by shore of Mazaruni River, Demerara, Dec 1890 = Isoetes clavata Weber, Berichte Deut. Bot. Gesellsch. 52: 121 (1934) = Isoetes goebelii H. Christ, ?=  I. triangula

Isoetes ovata V. Vareschi, 1968, Fl. Venezuela 1 (1): 110 (non Pfeiffer, 1922). [I. triangula Weber, 1922 (= ? I. schinzii Fuchs, 1960 nom. nud.) = Isoetes triangula Weber, 1922

Isoetes pacifica Svenson 1944, Amer. Fern. J. 34 (4): 123 = I. panamensis

Isoetes pallida Hickey, 1988, Amer. Fern J. 78(1): 35 - Mexico (Oaxaca)

Isoetes palmeri H.P.Fuchs, 1981, Proc. Kon. Ned. Akad. Wetensch. C 84(2): 168. 2n = 44 [Isoetes cleefii Fuchs-Eckert, 1981, Proc. koninkl. nederl. Akad. Wetensch. 84 (2): 177-179, ill. ?= I. boyacensis Fuchs in sched.). Type: Venezuela: Laguna Grande de Apartaderos, Mérida. Columbia: Cundinamarca: Páramo de Sumapaz: Laguna Gubernador, 3815 m, S San Juan (I. cleefii); Boyacá, Cauca, Arauca, Meta, in permanent pools and lakes, 3340-4435  m; Ecuador: Chimborazo (?= I. killipii Morton, 1945 - Colombia: Boyacá, Meta, Nariño, Santandér).

Isoetes panamensis Maxon & C.V. Morton, 1939, Ann. Missouri Bot. Garden 26(4): 272. 2n = 44 [Isoetes pacifica Svenson, American Fern Journal 34: 123 (1944); Isoetes savannarum L.D.Gómez-Pignataro, Phytologia 49(4): 339 (1981)] - Costa Rica: Guanacaste, inmediationem Crucis, prope loco Tanques dicto, ca. 200 m, Panama, Guatemala, Colombia, Ecuador: Guyabas, at base of Chanduy Mts; E Peru: Tumbes; (up to 1500 m). Ill.  microspores  MUSSELMAN, 2002.

Isoetes panchananii Pant & Srivastava, 1962, The genus Isoetes in India. Proc. Nat. Inst. Sci. India, vol. 28, part B: 242-280 - India: Madhya Pradesh: Rewa. Ill. loc. cit. 266, 267, map: 277 = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes panchganiensis Srivastava, Pant & Shukla, 1993, Amer. Fern J. 83(4): 107 (I. p. var. kemangundiensis) - Maharashtra, Karnataka. I. p. subsp. kemangundiensis Srivastava, Pant & Shukla - Karnataka = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes pantii Goswani & Arya, 1970, Journ. Indian Bot. Soc. 49: 30, 1968 (1970), - Madhya Pradesh: Narsinghghar, 80 km from Bhopal. Ill. loc. cit.: 36 = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes paraguayensis, Fuchs, 1960, nom. nud. (I. gardneriana sensi Hassler, 1928, Trabaj. Inst. Bot. Farm., Buenos-Aires 45: 98). Locus class.: Paraguay: Caaguazú: Guarapi; also: Pilcomayo, President Hayes, Misiones and dept. Central. = I. ekmannii

Isoetes paranaensis Fuchs, 1960, nom. nud. Locus class.: Argentina: Buenos Aires: Correa: delta del Paraná (?= I.  smithii)

Isoetes parryi = I. bolanderi

Isoetes parvula Hickey, 1994, Fieldiana, Bot. 34: 92 - Peru: Ayacucho. Known from type only.

Isoetes paupercula (Engelm.) A.A. Eaton = I. occidentalis

Isoetes pedersenii Fuchs-Eckert ex Hickey, 1980, nom. nud. (I. gardeneriana sensu Pedersen in sched.). Locus class.: Argentina: Corrientes: Depto Mburucuyá: Estancia Santa Maria. ?Bolivia (= I. paraguayensis =?)

Isoetes perralderiana Milde, 1867, Fil. Eur. Atlant., p. 282 = I. velata subsp. perralderiana

Isoetes perreymondi = I. velata

Isoetes perrieriana Iversen, 1932, Dansk Botanisk Arkiv 7: 200-201 - Madagascar. 

Isoetes peruviana U.Weber = I. lechleri

Isoetes phaeospora Durieu = I. tripus

Isoetes philippinensis Merrill & Perry, 1940, Am. Fern Journ. 30: 19, fig. - Philippines: Mindanao. Map Marsden, 1979: 342

Isoëtes phrygia Hausskn.,1899, Mitth. Thüring. Bot. Vereins, 13: 72 = I. histrix

Isoetes pichichensis Fuchs-Eckert, 1981, nom. nud. Locus class.: Ecuador: Pichincha, 4000 m. Also: Ecuador: Chimborazo, 3700 m, Volcán Chiles, 3900-4050 m, Páramo El Angel, 3750-3850, Colombia: Nariño, 4000-4100 m ?= I. novogranadensis, ?= I. karstenii

Isoetes piedmontana (N. Pfeiffer) C. Reed = I. melanopoda

Isoetes pilularioides Fuchs-Eckert, 1981, nom. nud. Locus class.: Brazil: Porto Alegre: Vila Manresa, ca 250 m.

Isoetes piperi A.A.Eaton = I. occidentalis

Isoetes pitotii Alston, 1956, Bol. Soc. Broter., 2A, 30: 15 - Mali, Guinea.

Isoetes precocia R.L. Small & Hickey, 2001, Fern J. 91(2): 61-62, figs. 12-13, 18-19. 2n = 22 [I. socia sensu Fuchs-Eckert, 1982, pro parte, Proc. Kon. Ned. Wetensch. C 85 (2): 256, non A. Braun] (near I. karstenii) - Venezuela: Sierra de St. Domingo, Páramo de Mucubají, road to Laguna Negra, ca. 3500 m; Columbia: Boyacá: Sierra Nevada del Cocuy, Alto Valle Lagunillas NE Laguna Pintada, 4390 m, and Boquerón de Cusiri, 4310 m; Páramo de la Sarna entre Sogamoso y Vado Hondo, 3330 m;  Cundinamarca: Páramo de Sumapaz, Chisaca SW Laguna Larga, 3620- 3700 m; Páramo de Cruz Verde, 3200 m; Meta: Páramo de Sumapaz, Cerro nevado del Sumapaz, 3725 m, in ephemeral pools.

Isoetes pringlei Underw., 1890, Zoë 1: 98 - Mexico (Jalisco, Mexico State).

Isoetes prototypus D.M. Britton, 1991, Canad. J. Bot. 69: 278 - E Canada (Nova Scotia, New Brunswick), northeastern USA (Maine).

Isoetes pseudojaponica M. Takamiya, Watan. & Ono, 1998 - Japan.

Isoetes pusilla C.R.Marsden & R.J. Chinnock, 1993, Muelleria 8(1): 57 - Australia: Victoria. Map Marsden, 1979: 334.

Isoetes pygmaea Engelm. = I. bolanderi

Isoetes quilindanensis Fuchs-Eckert, 1981, nom. nud.. Locus class.: Ecuador: Napo: Cordillera Oriental, small streams around Laguna Verdecocha = prob. I. novogranadensis

Isoetes rajasthanensis Gena & Bhardwaja, 1984. J. Bombay Nat. Hist. Soc. 81: 165-168 - India: Rajasthan: Mt. Abu, Mainal. Ill. loc. cit.:165 = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes ramboi forma latifolia Herter = I. smithii

Isoetes ramboi G. Herter, 1949, Rev. Sudamer. Bot. 8 (1-2): 19-25 - Brazil: Rio Grande do Sul. Ill. Fuchs, 1986: 23; MUSSELMAN, 2002 (microspores)= I. smithii

Isoetes rauhii Fuchs, 1960, nom. nud. Locus class.: Peru: Ancash: Cordillera Raura, 4500 m. Also: Cajamarca: Laguna Las Compuertas, Junín: prov. Yauli, 4800 m, Bolivia: Cochabamba: prov. Ayopaya, 3060 m.

Isoetes regulensis = I. velata

Isoetes reitzii Fuchs-Eckert, 1979, nom. nud. pro syn. I. brasiliensis. Locus class.: Brasil: São Leopoldo: Capao da Lagoa, 30 m = I. brasiliensis

Isoetes reticulata Gena & Bhardwaja, 1983, J. Bombay Nat. Hist. Soc. 81: 165-168 - India: Rajasthan: Atru, Ajmer, Jhalawar. Ill. loc. cit.:165 = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes rhodesiana Alston, 1956, Bol. Soc. Broter., ser. 2, 30: 17 - Zimbabwe: Bongola.

Isoetes rimbachiana Fuchs-Eckert, Proc. Kon. Ned. Akad. Wetensch. C 85(2): 226 (1982), Proc. koninkl. nederl. Akad. Wetensch. 85 (2): 226. Locus class.: Ecuador: Mount Chimborazo; Pinchincha, Napo = I. karstenii var. anomala

Isoetes riparia Engelm. ex A. Br., 1846, Flora (Regensb. Bot. Zeit.) 29: 178 [Calamaria riparia (Engelm. ex A. Br.) Kuntze, Rev. Gen. Pl. 2: 828 (1891) = Isoetes foveolata var. plenospora Eaton, Rhodora 5: 280 (1903; Isoetes riparia var. riparia Proctor, Amer. Fern Journ. 39: 119 (1949); Isoetes riparia var. typica Proctor. Amer. Fern Journ. 39: 119-121 (1949); Isoetes riparia var. canadensis Engelm. ex N.Pfeiffer, Trans. St. Louis Acad. Sci. 4: 383 (1882) Isoetes canadensis (Engelm.) A.A. Eaton, Proceedings of the United States National Museum 23(1226): 650 (1901); Isoetes dodgei Eaton, Fern Bull. 6: 6 (1898) Isoetes riparia forma canadensis (Engelm.) Proctor.; Isoetes riparia var. robbinsii forma canadensis (Engelm. ex Pfeiffer) Proctor, Amer. Fern Journ. 39: 119 (1949), Isoetes riparia forma saccharata (Engelm.) Clute, Fern. Bull. 16: 57 (1908); Isoetes saccharata forma palmeri Eaton, Fern Bull. 11: 25 (1903); Isoetes saccharata var. palmeri A.A. Eaton, Proceedings of the Biological Society of Washington 14(10): 49 (1901); Isoetes saccharata var. palmeri (A.A.Eaton) Proctor, Amer. Fern Journ. 39: 119 (1949); Isoetes amesii Eaton, Gilbert, Fern Bull. 11: 103, 1903; Isoetes saccharata var. amesii (A.A.Eaton) Proctor, Amer. Fern Journ. 39: 119 (1949); Isoetes saccharata var. amesii A.A. Eaton, Rhodora 5(59): 278-279 (1903), Isoetes canadensis var. amesii (Eaton) Clute, Fern Bull. 16: 54 (1908);  Calamaria saccharata (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes saccharata Engelm., Gray's Manual, ed. 5, 676 (1867); Isoetes riparia forma saccharata (Engelm. ex A. Gray) Proctor, Isoetes riparia var. palmeri forma saccharata (Engelm.) Proctor, Amer. Fern Journ. 39: 119 (1949); Isoetes saccharata var. reticulata A.A.Eaton) Proctor, Amer. Fern Journ. 39: 119 (1949) Isoetes saccharata forma reticulata Eaton, Fern Bull. 11: 25 (1903);  Isoetes saccharata var. reticulata Eaton; Proc. Biol. Soc. Wash. 14: 49 (1901); Isoetes riparia var. robbinsii (Eaton) Proctor, Amer. Fern Journ. 39: 119 (1949); Isoetes canadensis var. robbinsii A. A. Eaton, Rhodora 5: 279 (1903); Isoetes dodgei var. robbinsii Eaton, Rhodora 10: 42 (1908); Isoetes chapmanii var. robbinsii (Eaton) Broun, Index N. Amer. Ferns 199 (1938); Isoetes hyemalis D.F. Brunton & Britton, 1994] - E Canada (S Quebec, S Ontario), Eastern USA (Connecticut, Delaware, Florida, Maine, Maryland, Massachusetts, New Hampshire, New Jersey, New York, North Carolina, Pennsylvania, Rhode Island, South Carolina, Vermont, Virginia, West Virginia, Carolina and Forida - Map Fl. N Amer.: 71. Ill. Fassett: 46, AWP 1: 26, Taylor & Hickey, 1992: 618, Crow & Hellquist 1: 26.

Isoetes roncadorensis Fuchs-Eckert, 1981, nom. nud. Locus class.: Brazil: Mato Grosso: Serra do Roncador, ca. 450 m (= ?

Isoetes roraimensis Fuchs-Eckert, 1981, nom. nud. Locus class: Venezuela: Bolivar: Mount Roraima, 2700-2740 m. (very similar to I. killipii Morton, 1945 - Fuchs-Eckert, 1982: 248) prob. = I. killipii

Isoetes rosselii Fuchs-Eckert, 1964, nom. nud. Locus class.: Brasil: Bahia: Rio Preto. Map Fuchs, 1982 (I. van-nielii).

Isoetes saccharata Engelm. in A. Gray (I. hyemalis) = I. riparia

Isoetes sahayadrii Mahabale, 1938, Current Science, vol. 7 (2): 62-63. - Karnataka and Maharashtra: Panchgani = I. coromandelina

Isoetes sampathkumaranii Rao, 1944, Current Science, vol. 13 (11): 286-287 - India: Bangalore, Karnataka. Ill. p. 274 and map: 277 in Pant & Srivastava, 1962 = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes saracochensis Hickey, 1994, Fieldiana 34: 94 (I. saracochensis Fuchs, 1960, nom. nud.). Locus class.: Perú: Laguna Saracocha, 4200 m; also: Lagunillas, 4380 m, Laguna Saytococha, 4350 m.

Isoetes savannarum L.D.Gómez = I. panamensis

Isoetes savatieri Franchet, 1884, Bull. Soc. Bot. Fr. 31: 395 (aff. I. setacea) [Calamaria savatieri (Franchet) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes lechleri var. savatieri (Franchet) L.D. Gómez, Brenesia 18: 5 (1980; Isoetes valdiviensis Fuchs, 1960, Isoetes savatieri subsp. valdiviensis Fuchs-Eckert (1992) nom. nud.; I. meyeri Fuchs-Eckert, 1982 (nom. nud.); I. chilensis Fuchs-Eckert, 1981 (nom. nud.)]. (However see I. chubutiana Hickey, Macluf & Taylor, 2003) - Andes S Argentina: Neuquén, lake Nahuel Huapi etc., Rio Negro, Chubut, Santa Cruz, Tierra del Fuego, and Chile 770-1460 m (Arauco 37° S to Patagonia, Puerto Bono, Tierra del Fuego). Map Fuchs-Eckert, 1982; ill.  microspores  MUSSELMAN, 2002.

Isoetes savatieri subsp. valdiviensis H.P. Fuchs-Eckert, 1992: 140 (nom. nud.) = I. savatieri

Isoetes schinzii Fuchs, 1960, nom. nud. - British Guyana: Essequibo; Guyane Française (= I. ovata ?= I. triangula)

Isoetes schweinfurthii A. Braun, 1880, Journal of Botany, British and Foreign 18: 108 [Calamaria schweinfurthii (A. Braun) Kuntze, Revisio Generum Plantarum 2: 828 (1891) 1893; Isoetes alstonii C. Reed & Verdc., Kirkia 5: 19-20, 1965; Isoetes kersii Wanntorp, Svensk Bot. Tidskr. 64: 146, f 3, 4, t. 1, 3, 1970; Isoetes rhodesiana Alston, 1956] - Morocco, Senegal, Ivory Coast, Central African Republic, Sudan, Chad, Zambia, Zimbabwe, Tanzania, Mozambique, Botswana, Transvaal, Namibia; Madagascar: Mahajanga, Betafo. Ill. COOK, 2004: 34

Isoetes sehnemii H.P. Fuchs, 1986, Fl. Illustr. Catar. 1 (Isoetaceas): 38 - Brazil (Santa Catarina, Rio Grande do Sul: Vacaria, 400-450 m).

Isoetes semilocularis J.E. Smith, Rees, 1819, Cyclopaedia 19: 150 (1819) - Distribution unknown = Isoetes coromandelina sensu Roxb. ex Griff., Calcutta Journ. Nat. Hist. 4: 470 (1844).

Isoetes setacea Lam., 1789, Encyclopédie Méthodique, Botanique 3: 314 (nom. ambig. fide Rothmaler, 1944). 

Isoetes setacea Lam., Encyclopédie Méthodique, Botanique 3: 314, 1789 2n = 22 [Calamaria setacea (Bosc ex Delile non Lam.) Kuntze, Revisio Generum Plantarum 2: 828 (1891), Isoetes setacea Bosc ex Delile, 1827, Mém. Mus. hist. nat. Paris 14: 100-119 (nom ambig. fide Rothmaler; I. echinospora Durieu, 1861, Isoetes echinospora mult. auct. Map Hultén & Fries, 1: 5, TAYLOR et al., p. 68. Ill. Prelli: 83; BERTHET & LECOCQ, 1977: 335, 355, 356. Isoetes setacea Lam. subsp. setacea - W Palaearctic S to the Pyrenees, N Italy. . Isoetes setacea subsp. asiatica (Isoetes echinospora subsp. asiatica (Makino) A.Löve, Amer. Fern J. 52: 121. 1962  [Isoetes asiatica (Makino) Makino, Bot. Mag. Tokyo 28: 184 (1914); Isoetes echinospora subsp. asiatica (Makino) A. Löve, Isoetes echinospora var. asiatica Makino, Bot. Mag. (Tokyo) 18: 129. 1904 (Iversen, Dansk Bot. Ark. 5 (23): 2.1928), Isoetes muricata Durieu subsp. asiatica (Makino) H.P.Fuchs, Proc. Kon. Ned. Akad. Wetensch. C 85 (2): 239 (1982)  - Mongolia, S Kuril Isl., Sakhalin, Kamchatka, Japan, ?China, Japan and Korea to Kamtchatka. Map Hultén & Fries 1: 5; map and ill. Kadono 8, 10. Isoetes setacea subsp. braunii [Isoetes braunii Durieu, Bull. Soc. Bot. France 11: 101. 1864 Isoetes echinospora var. braunii (Durieu) Engelm. in A.Gray, Isoetes muricata var. braunii (Durieu) C.F.Reed.  Isoetes tenella var. braunii (Durieu) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 73, 1944; I. muricata Dur., Isoetes echinospora subsp. muricata (Durieu) A. Löve & D. Löve,  Isoetes echinospora var. muricata (Durieu) Engelm. in A.Gray, Manual (Gray), ed. 5 676, 1867, I. echinospora subsp. muricata (Durieu) A.Löve & D. Löve, Isoetes echinospora f. astoma Iversen, Bot. Tidsskr. 40: 129, 1928, Isoetes braunii Durieu f. astoma (Iversen) M.Broun, Index No. Amer. Ferns 99, 1938, Isoetes tenella f. astoma (Iversen) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 72, 1944, Isoetes braunii Durieu f. oligostoma (Iversen) M.Broun, Index No. Amer. Ferns 99, 1938,  Isoetes tenella f. oligostoma Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 73, 1944, Isoetes tenella var. oligostoma (Iversen) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 72, 1944, Isoetes braunii Durieu f. polystoma (Iversen) M.Broun, Index No. Amer. Ferns 99, 1938 Isoetes tenella f. polystoma Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 73, 1944;  Isoetes echinospora f. polystoma Iversen, Bot. Tidsskr. 40: 128. 1928 Isoetes tenella forma robusta (Engelm.) Rothm. Isoetes braunii Durieu f. robusta N.Pfeiff., Ann. Missouri Bot. Gard. 9: 173, 1922  Isoetes echinospora var. robusta Engelm., Isoetes echinospora f. robusta (Engelm.) Clute, Fern Allies 258, 1905 . Isoetes muricata Durieu f. robusta (Engelm.) C.F.Reed, Amer. Fern J. 35: 82, 1945,  Isoetes tenella f. robusta (Engelm.) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 73, 1944, Isoetes echinospora var. robusta Engelm., Trans. Acad. Sci. St. Louis 4: 380 (reprint p. 23), 1882,  Isoetes echinospora f. boottii (Engelm.) Clute, Fern Allies 258, 1905 Isoetes echinospora var. boottii Engelm., Isoetes tenella forma oligostoma Rothm., Isoetes tenella var. oligostoma (Iversen) Rothm., Isoetes braunii forma oligostoma (Iversen) M.Broun., Isoetes echinospora f. oligostoma Iversen, Bot. Tidsskr. 40: 129, 1928, Isoetes echinospora var. brittoni Cockerell, Isoetes echinospora [infrasp.unranked] brittonii Cockerell, Muhlenbergia 3: 9, 1907. Isoetes echinospora subf. lativelata Iversen, Bot. Tidsskr. 40: 129 fig. d. 1928,  Isoetes tenella f. lativelata (Iversen) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 72, 1944, Isoetes echinospora subf. angustivelata Iversen, Bot. Tidsskr. 40: 128, fig. a-c. 1928,  Isoetes tenella f. iversenii Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 73, 1944, I. hesperina. see also synonymy sub I. echinospora;  see also synonymy sub I. echinospora - Alaska, Canada (Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, Labrador, Nunavut, NW. Territories, Nova Scotia, Ontario, Prince Edward Isl., Quebec, Saskatchewan, Yukon), NW USA, NE-USA (California, Colorado, Connecticut, Idaho, Maine, Massachusetts, Michigan, Minnesota, Rhode Island, Utah, Vermont, Washington, Wisconsin), Barbados. Map Fl. N Amer.: 68; Muenscher: 343; ill. Crow & Hellquist 1: 26; Fassett: 46; Muenscher: 341.

Isoëtes setacea auct., sensu Jovet & R. Vilmorin, sensu A.C. Jermy [1964, Fl. Eur., 1: 5], sensu A. Lawalrée & M. Guinochet in 5 [1973, Fl. Fr., 3: 134, n° 110] = I. setacea Lam., 1789.

Isoetes setacea Bertol. = I. velata

Isoetes setacea Bory = I. velata

Isoetes setacea forma submersa Glück = I. velata

Isoetes setacea forma terrestris Glück = I. velata

Isoetes setacea Moris & De Notaris = I. histrix

Isoetes setacea var. delilei Bory, 1844 = I. velata (in Pfeiffer, 1922: 119).

Isoetes setacea var. perreymondi = I. velata

Isoëtes sicula Todaro,1866, Giorn. Sci. Nat. Econ. Palermo, 1: 241 = I. subinermis

Isoetes sicula Tod., Syn. Pl. Acot. Vasc. Sic. 251 (1866); I. histrix subsp. sicula (Todaro) P. Fournier = I. subinermis

Isoetes sinensis Palmer, Amer. Fern J. 17: 112 (1927) (I. japonica subsp. sinensis) (2n = 44) - Japan, China: Anhui, Jiangsu, Zhejiang, alt. 130-300 m. (?= I. japonica subsp. sinensis)

Isoetes sinensis var. coreana (Y.H. Chung & H.K. Choi) M. Takamiya & al. = I. japonica subsp. sinensis

Isoetes smithii H.P. Fuchs, 1986, Fl. Illustr. Catar. 1 (Isoetaceas): 17 [Calamaria martii (A. Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes martii A. Br., ex Kuhn, in Martius, Fl. Bras. 1 (2): 646, t. 78 (1884); Isoetes ramboi Herter, Rev. Sudamer. Bot. 8 (1-2): 19-25 (1949)]; Isoetes ramboi forma latifolia Herter; Isoetes weberi Herter ex Weber, Hedwigia 63: 255-256, fig. 44-45 (1922); (?I. kriegerii - Minas Gerais; ?I. herteri - Uruguay: Dept. de Rocha: Matavillas) - Brazil: Serra do Itatiaia, in small stream (type); Minas Gerais, Rio Grande do Sul: Santa Catarina; Uruguay. Map FUCHS, 1986: 20, ill.: 14; ill.   MUSSELMAN, 2002 (microspores).

Isoetes socia A.Braun, 1862, Verh. Bot. Ver. Brandenbg. 3-4: 332 (sectio Laeves - Fuchs, 1982) = I. lechleri

Isoetes spannagelii Fuchs-Eckert, 1986, Fl. Illustr. Catar. 1 (Isoetaceae): 33 - Brazil: Rio Grande do Sul: SE Santa Catarina: Lages, Vacas Cordas. Map Fuchs, 1986: 36, ill.: 34.

Isoetes spinulospora Jermy, apud Anthony & Schelpe, 1982, Contr. Bolus Herb. 10: 151. Known only from locus class.: Equatorial Guinea, Fernando Po: Bioko Isl.: Lagoa Biao.

Isoetes stellenbossiensis A.V. Duthie, 1929, Trans. R. Soc. S. Afr. 17: 328 - SW Cape Prov. Ill. COOK, 2004: 34

Isoetes stephansenii Duthie, 1929 - S Africa.

Isoetes stevensii Croft, 1980, Blumea 26 (1): 184 - Papua New Guinea: Mt. Giluwe, Mt. Sarawaket, 3400-3600 m. Map CROFT, 1980: 182, ill.: 187.

Isoetes steyermarkii Fuchs, 1960, nom. nud. Locus class.: Venezuela: Mérida, near El Aquila, 3930 m. Also: Mérida, 3400, 3700 m, Colombia: Boyacá, Cundinamarca, Cauca, Huila-Cauca, dept. del Valle, 3530-4000 m = Isoetes karstenii var. karstenii

Isoetes storkii T.C. Palmer, 1932, American Fern Journal 22(4): 136 [Isoetes montana T. C. Palmer, American Fern Journal 21(4): 134 (1931) [non Isoetes montana U. Weber (1922); Isoetes tryoniana L.D. Gómez, Revista de Biología Tropical 17(1): 108, f. 8 (1972)] - Costa Rica (Cordillera de Talamanca, 3300 m, San José: Dota Copey, Cerro vueltas), Panama.

Isoetes stuartii A.Braun 1868 = I. muelleri

Isoetes subinermis (Durieu) Cesca & Peruzzi, 2001, Flora Medit. 11: 303-309; 2n = 22 . Cephaloceraton gymnocarpum Genn., Comment. Critt. Ital. l (3): 113 (1862); Isoetes gymnocarpa (Genn.) Braun; Isoetes sicula Tod., Syn. pl. acot. vasc. Sic.: 46, 1866; Isoetes sicula Tod. ex Ces. & DeNot. apud Genn., Comm. Soc. Critt. lt. 1 (2): 103 (1861); Isoetes histrix subsp. sicula (Tod) Fournier, Quatre fl. de France 6, 1934 ; Isoetes histrix Bory var. subinermis Durieu; Cephaloceraton hystrix var subinermis (Durieu) Genn., Comment. Critt. Ital. 1 (3): 112 (1862); Isoetes hystrix subinermis Gand., Fl. Eur. 27: 197 (1891); Isoetes histrix forma subinermis Durieu, Bull. Soc. Bot. France 8: 164, 186 (1861); Isoetes histrix var. subinermis Dur., Cesati, Bull. Soc. bot. France 8: 164, Comp. Fl. lt., 25 (1867), et Motel. & Vendr., Actes Soc. Linn. Bord. 36: 4009, t. 17, f. 1-5 (1883);   Isoetes histrix β subinermis (Durieu) Fiori, Nuova Fl. analitica d'Italia: 144, 1926; Isoetes histrix var. subinermis forma submersa Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 20, 1936; Isoetes histrix var. desquamata A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec (1863): 617 (1864);  Isoetes histrix var. desquamata A. Br. forma subinermis (Durieu) Fiori, Fl. Ital. Crypt. 5: 424-426, 1943; Isoetes histrix var. desquamata A. Br., Batt. & Trab., Fl. Alg. & Tunis. 407 (1902); Isoetes histrix var. desquamata forma longispina (A. Br.) Fiori, Fl. Ital. Crypt., pars V: 425, f. 154 (1943); Isoetes histrix forma desquamata (a) solitaria A. Br., Monatsber. K. Akad. Berlin, Sitzung 7 Dec. (1863): 618 (1864); Isoetes histrix forma desquamata (b) caespitosa A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec. (1863): 618 (1864); Isoetes histrix forma desquamata (b) caespitosa brevispina A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec. (1863): 618 (1864); Isoetes histrix forma desquamata (b) caespitosa longispina A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec. (1863): 618 (1864); Isoetes histrix forma desquamata A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec (1863): 617 (1864); Isoetes histrix var. desquamata f. (b) subinermis Dur., Bull. Soc. Bot. Fr. 8: 164 (nomen) (1861)  Isoetes histrix f. desquamata (b) caespitosa subinermis A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec. (1863): 618 (1864)].


Isoetes suksdorfii Baker = I. nuttallii

Isoetes taiwanensis De Vol, 1972, Taiwania 17(1): 1-7, f. 1. 2n = 22 - Taiwan: Taipei County. Map Marsden, 1979: 342.

Isoetes tasmanica Durieu, 1864 = both I. elatior and I. humilior

Isoetes tegetiformans P.M. Rury, 1978, Amer. Fern J. 68(4): 100 - N Georgia. Ill. Fl. N Amer.: 71.

Isoetes tegulensis forma submersa = I. velata  subsp. tegulensis

Isoetes tegulensis forma terrestris = I. velata subsp. tegulensis

Isoetes tegulensis Gennari, 1861, Comment. Soc. crittog. ital. 1 (2): 106 (September), nomen illeg. = I. velata

Isoëtes tenella Léman ex Desv., 1827, Mém. Soc. Linn. Paris, 6 (2): 179 ?= I. setacea Lam. (I. echinospora)

Isoetes tennesseensis Luebke & Budke, 2003 (2n = 88) - USA: SE Tennessee: Hiwassee River.

Isoetes tenuifolia Jermy apud J. Hall, 1971, Bot. J. Linn. Soc. 64: 118 - Known only from type locality: Ghana: Accra Plains: Nungua Farm; ? Kenya.

Isoetes tenuissima Boreau, 1850, Bull. Soc. industr. Angers 21: 269 (Etang de Ris-Chauvron) = I. velata subsp. tenuissima

Isoetes ticlioensis Fuchs-Eckert, 1981, nom. nud. Locus class.: Perú: Dept. Lima: Ticlio pass, 4840 m. Known only from vicinity of Ticlio Pass.

Isoetes tiguliana Gennari, 1861, Comment. Soc. crittog. ital. 1 (1): 42 (February) = I. velata

Isoetes toximontana L.J. Musselman & J.P. Roux, 2002, Novon 12: 504-507 - S Africa: northern Cape: Gifberg. Ill. COOK, 2004: 34.

Isoetes transvaalensis Jermy & Schelpe, 1982, Contr. Bolus Herb. 10: 150 - Transvaal, Oranje Free State, Natal, Lesotho. Ill. COOK, 2004: 35.

Isoetes triangula U.Weber, 1922, Hedwigia 63: 253, f. 32, 33 [Isoetes ovata sensu Vareschi, 1968, Fl. Venezuela 1 (1): 110; I. martii sensu Ule in sched.; Isoetes guyanae U. Weber, nom. nud. (Fuchs-Eckert, 1982) = I. clavata Weber, 1934,  I. goebelii Christ ined., non Weber, 1922] - Locus class.: Brazil: Amazonas: Serra do Mel, Rio Branco, Surumu; SE Venezuela; British Guyana: Essequibo British Guyana: Rio Mazaruni, Baracarra, Demerara(I. ovata Pfeiffer, 1922), Guyane française: Tonate (I. schinzii Fuchs, ined.). Ill. Tryon, 1982: 833.

Isoetes tridentata =I. durieui

Isoetes tripus A. Br. ex Herb. Vindob. & Hook., 1863, Monatsbericht der Königlichen Akademie der Wissenschaften [Calamaria tripus (A. Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes phaeospora Dur., Bull. Soc. Bot. Fr. 11: 103 (1864)] - SW Western Australia. Map Marsden, 1979: 336.

Isoetes triquetra A. Braun, 1862, Verh. Bot. Vereins Prov. Brandenburg 4: 332 (nearly identical to I. andina - Hickey) [Calamaria triquetra, Isoetes lechleri var. triquetra (A.Br.) L.D. Gómez] - Andes of Ecuador and Perú: Sachapata. Cf. Hickey, R.J., 1986, Taxon 35: 243-246; I. triquetra var. dissimilis G. Rodríguez, 1955, Bol. Mus. Cienc. nat. Caracas 1 (2): 59 - Venezuela: Mérida: Laguna de Mucubají, 3500 m. =  I. andina

Isoetes truncata Clute = I. maritima

Isoetes tryoniana Gómez-Pignataro, 1970, Rev. Biol. trop. 17 (1): 108-117. Known only from type locality: Costa Rica: Cordillera de Talamanca, 3300 m.

Isoetes tuberculata Gena & Bhardwaja, 1983. J. Bombay Nat. Hist. Soc. 81: 165-168 - India: Rajasthan. Ill. loc. cit.:165 = I. coromandelina fide Fraser-Jenkins, 1997.

Isoetes tuckermanii A. Braun ex Engelm., 1867, A. Gray, Manual ed. 5: 676 [Calamaria tuckermani (A. Braun ex Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes tuckermani forma borealis (Eaton) Clute, Fern Allies, 260 (1905); Isoetes tuckermani var. borealis A.A.Eaton, Fern Bull. 6: 7 (1898); Isoetes tuckermani var. harveyi Clute] - E Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario, Quebec), northeastern USA (Connecticut, Maine, Massachusetts, New Hampshire, New Jersey, New York, Rhode Island, Vermont). Map Fl. N. Amer.: 70. Ill. Fassett: 46; Muenscher: 341; AWP 1: 26; Crow & Hellquist 1: 26.

Isoetes tuckermannii var. harveyii = I. x harveyii

Isoetes tuckermannii var. heterospora = I. x heterospora

Isoetes tuckermannii x I. engelmannii (I. x foveolata)

Isoetes tuerckheimii Brause in Urban., 1912, Symb. Antillianae 7: 161 - Hispaniola (Haiti, up to 2200 m; Dominican Republic ).

Isoetes udupiensis P.D. Shukla, G.K. Srivastava,  S.K. Shukla & P.K. Rajagopal, 2005. Taxon, 54 (1): 109-116. - Peninsular India.

Isoetes ulei U.Weber, 1922, Hedwigia 63: 255, fig. 40-41 - Brazil: Rio de Janeiro.

Isoetes underwoodi L.F. Hend. = I. howellii

Isoetes unilocularis J.E. Smith (I. indica Pant & Srivastava non J.G. König ex J.E. Smith; I. mirzapurensis Pangrahi & Dixit) - India: Madhya Pradesh, Utar Pradesh, Tamil Nadu. = I. coromandelina

Isoetes urussuhuyensis Fuchs-Eckert, 1981, nom. nud. Locus class.: Brasil: Piauhy: Rio Parnahyba, Serra do Urrussuhuy. Also: Lagoa Boqueirão, S Piauhy.

Isoetes valdiviensis Fuchs, 1960 = I. savatieri

Isoetes valida (Engelm.) Clute, 1905, The Fern Allies 236 [Isoetes valida var. gravesii (A.A.Eaton) Clute, I. engelmannii var. valida Engelm., Isoetes engelmannii var. caroliniana A.A. Eaton, Fern Bulletin 8: 60 (1900)] - Eastern USA. Fl. N Amer.: 69. Cf. Amer. Fern J., 86: 16-25, 1996.

Isoetes van-nielii Fuchs, 1960, nom. nud. Locus class.: Brasil: Bahia: Rio Preto (I. rosselii, cf. I. graminifolia).

Isoëtes variabilis Le Grand, 1893, Notes Isoètes du Centre: 11-12 (nom. illeg.) = I. velata

Isoetes vaupesensis Fuchs-Eckert, 1981, nom. nud. ined. Locus class.: Colombia: Vaupés: Intica. = prob. I. triangula

Isoetes velata A. Braun in Bory de Saint-Vincent & Durieu de Maisonneuve, 1848, Expl. Sci. Algérie, Atlas: tab. 37, fig. 1. - Ill. C & K 1: 52, 57; Prelli: 83; BERTHET & LECOCQ, 1977: 345. Isoetes velata subsp. velata A. Braun, Bory & Durieu, 1849, Expl. Sci. Algérie, Atlas: tab. fig. 1. 2n = 22. Isoetes decipiens Bory, Flora 29: 719 (1846) deg. nom. prov.; Isoetes grammontii Bory 1841 in Milde, Fil. Eur. & Alt., Asiae Minores & Siberiae, 280 (1867) Isoetes variabilis Le Grand, Mém. Soc. Hist. Cher ser. 4, 1896: 11 (1896), nom. illeg.; Isoetes velata algeriensis Gdgr., Fl. Eur. 27: 195 (1891); Isoetes velata australis Gdgr., Fl. Eur. 27: 195 (1891); Isoetes velata barbara Gdgr. Fl. Eur. 27: 194 (1891); Isoetes velata divergens Gdgr. Fl. Eur. 27: 195 (1891); Isoetes velata forma algerica A. Br. ex Milde, Fil. Eur. & Atl., 282 (1867); Isoetes velata forma brevifolia A. Br. ex Motel & Vendr., Actes Soc. L. Bord. 36: 384 (1883); Isoetes velata subsp. typica fa. algerica A.Br., Monatsber. Akad. Wiss. Berlin, p. 605 (1863), Isoetes velata subsp. typica forma algerica (A. Br.) Maire & Weiller, Encycl. Biol. 33: 90 (1952) Isoetes velata subsp. typica forma brevifolia (A. Br.) Maire & Weiller, Encycl. Biol. 33: 90 (1952); Isoetes velata var. typica forma brevifolia A. Br., Monatsber. K. Akad. Wiss. Berlin, Sitzung 7 Dec (1863); Isoetes velata var. brevifolia A. Br., Batt. & Trabut, Fl. Alg. & Tunis., 407 (1902); Isoetes velata forma humilior Marcucci, Un. Itin. Crypt., No. 1-V (1866)Isoetes velata var. humilior A. Br. ex Mares & Vigineix, Pl. Vasc. des Iles Baléares, 322 (1890); Isoetes velata gallica Gdgr., Fl. Eur. 27: 194 (1891); Isoetes velata redux Gdgr., Fl. Eur. 27: 195 (1891); Isoetes velata robusta Gdgr., Fl. Eur. 27: 194 (1891); Isoetes velata subsp. typica (Fiori & Paol.) Maire & Weiller, Fl. Anal. Ital., p. 25 (1895);  Isoetes velata subsp. typica (Fiori & Paol.) Maire & Weiller, Encycl. Biol. 33: 90, f. 51 (1952); Isoetes velata var. typica Fiori & Paoletti, Fl. Anal. Ital. 1: 25 (1896); Isoetes velata forma lusitanica A. Br. ex Milde, Fil. Eur. & Atl., 282 (1867); Isoetes velata tenuicula Gdgr., Fl. Eur. 27: 195 (1891); Isoetes velata trinacria Gdgr., Fl. Eur. 27: 195 (1891); Isoetes velata var. brevifolia A. Br., Sitzungsber. Berlin. Akad. Wiss. (Monatsber. Preuss. Akad. Wiss. Berlin), 605 (1863); Isoetes velata var. brevifolia (A. Br.) Franchet, Bull. Soc. Bot. Fr. 31: 349 (1884); Isoetes velata var. normalis A. Br., Bory & Dur., Expl. Sci. Alg., t. 37, fig. 1 (1848)Isoetes perreymondi (Bory) Duval-Jouve, Bull. Soc. Bot. Fr. 16: 213 (1869); Isoetes adspersa var. perreymondii A. Br., Schoenefeld, Bull. Soc. Bot. Fr. 12: 261 (1865); Isoetes velata subsp. tenuissima var. perreymondii (A. Braun) P. Fourn., 1934, Quatre Fl. Fr.: 5 (comb. illeg.); Calamaria dubia (Genn.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes dubia Genn., Comment. Soc. Crittog. Ital. 1: 104 (1861); I. velata subsp. dubia (Genn.) Batt. & Trabut, Fl. Syn. 407 (1902); Isoetes dubia var. emersa Genn., Nuovo Giorn. Bot. Ital. 2: 96 (1870); I. dubia var. vaculosa; Isoetes capillacea Bory, Compt. Rend. Inst. France 23: 620 (1846); Isoetes lineolata Dur. ex Motel. & Vendr., Actes Soc. Linn. Bord. 36: 381 (1883); Isoetes setacea Bory, Compt. Rend. Inst. France 18: 1165 (1844); Isoetes setacea var. perreymondii Bory, Compt. Rend. Acad. Paris 18: 1165 (1844); Isoetes velata perreymondi (A. Br.), Franchet, Bull. Soc. Bot. Fr. 31: 349 (1884); Isoetes velata var. perreymondi (Bory) Franchet]Isoetes baetica Willk. & Lange, Prodr. Fl. Hispan. 1: 14 (1861); I. velata forma baetica (Willk.) Prada, Acta Bot. Malac. 8: 88 (1983); I. velata forma lereschii (Rchb. ex Leresche & Levier) Prada, Lazaroa 3: 33 (1981 publ. 1982); Isoetes boryana var. lereschii Reichb. apud Ler. & Lev., Deux Exc. Bot. Esp. 118 (1880). I. velata subsp. velata forma baetica (Willk.) Prada, 1983, Acta bot. malacitana 8: 88, et forma lereschii (Reich.) M.C. Prada, 1982, Lazaroa 3: 33 (Spain: Sierra de Gredos); Isoetes boryana gredensis Gdgr., Fl. Eur. 27: 195 (1891) Spain, Portugal,  Baleares, France, Corsica,  Italy, Algeria, Libya, Tunisia, Morocco. Isoetes velata subsp. longissima (Bory) Greuter & Burdet, 1984, Willdenowia 14(1): 37; Calamaria longissima (Bory & Dur.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes longissima Bory, Compt. Rend. Hebd. Séances Acad. Sci. 18: 1165 (1844), basion. (Algeria); Isoetes velata forma longissima (Bory & Dur.) N. Pfeiff., Ann. of the Missouri Bot. Garden 9(2): 120 (1922); Isoetes velata forma longissima A. Br. ex Milde, Fil. Eur. & Atl. 282 (1867); Isoetes velata subsp. typica forma longissima (A. Br.) Maire, Encycl. Biol. 33: 90 (1952); Isoetes velata forma longissima Maire, Fl. Afr. Nord 91 (1952);  Isoetes velata var. longissima (A. Br.) Bory & Dur., Explor. Sci. Alg., t. 37, fig. 2 (1848); Isoetes velata var. longissima (A. Br.) Batt. & Trabut, Fl. Alg. et Tunisie, 407 (1902); Isoetes velata subsp. typica forma longissima A. Br., Expl. Scient. Alg., tab. 37, fig. 2 (1848), pro var.] - Algeria: El Kala, lac Houbeïra, lac Freitis, lac des Canards chez les Senhadja. Isoetes velata subsp. adspersa (A. Braun) Batt. & Trabut, Fl. Algérie Tunisie: 407 (1905); Calamaria adspersa (A. Br. in Bory & Dur.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes adspersa A. Braun, Bory, Expl. Sci. Algérie, Atlas: tab. 37, rig. 3 (1849); Isoetes aspersum Aschers. & Graebn., Syn. Mitteleur. Fl. 1: 169 Hauptreg. 23 (1897); I. velata subsp. tenuissima subsp. adspersa (A. Braun) Douin in Bonnier, 1934, Fl. Compl. Fr., 12: 118 (comb. illeg.); Isoetes velata proles adspersa (A. Braun) Rouy,1913, Fl. Fr., 14: 478 (comb. illeg.);  I. velata subsp. tenuissima [var.] adspersa (A. Braun) P. Fourn., 1934, Quatre Fl. Fr.: 5 (comb. illeg.); Isoetes velata ssp. perralderiana (Milde) Batt. & Trabut, 1905, Fl. Algérie Tunisie: 407; Isoetes perralderiana Dur. & Le Tourn. (Kralik), Pl. Alg. Exsicc. 157 (1862); Isoetes perralderiana Milde, Fil. Eur.: 282 (1867), basion.; Isoetes velata subsp. perralderiana (Dur. & Le Tourn.) Trabut, Fl. Syn. 407 (1902); Calamaria perralderiana (Dur. & Le Tourn., Milde) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes velata var. perralderiana (Dur. & Le Tourn.) Pfeiffer, Ann. Mo. Bot. Gard. 9: 121 (1922). Isoetes velata subsp. intermedia (Trabut) Maire & Weiller, Maire, Fl. Afrique N. 1: 89 (1952); Isoetes velata forma intermedia A. Br., Motel. & Vendr., Actes Soc. L. Bordeaux 36: 384 (1883); Isoetes velata subsp. intermedia forma immaculata Maire, Encycl. Biol. 33: 90 (1952); Isoetes velata tax. infrasp. intermedia Trabut, Batt. & Trabut, Fl. Algérie Tunisie: 407 (1905); Isoetes velata var. intermedia A. Br., Bory & Dur., Explor. Sci. Alg. (1848)]. Isoetes velata subsp. asturicensis (Lainz) Rivas Martinez & Prada, Lazaroa 2: 327 (1980), locus class.: Asturias: Cueto de Arbas; Isoetes boryana subsp. asturicensis Lainz, Bol. Inst. Estud. Asturianos, Supl. Ci. 15: 6 (1970); Isoetes velatum var. asturicense (M. Laínz) E. Rico & X. Giraldez, Anales Jard. Bot. Madrid 46(2): 585 (1989 publ. 1990). Ill. ROMERO et al., 2004. Isoetes velata subsp. tegulensis (Genn.) Trabut, Fl. Syn. 407 (1902); Calamaria tegulensis (Genn.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); I. tegulensis Gennari Comment. Critt. Ital. 1 (1d): 42, 1861, nom. illeg.;  I. velata subsp. tegulensis Gennari, loc. cit. 1 (2): 106, 1861, I. v. subsp. tegulensis Batt. & Trabut, Fl. Algérie Tunisie: 407 (1861); Isoetes velata var. tegulensis (Genn.) Fiori & Paoletti, Fl. Anal. Ital. 1: 25 (1896); Isoetes velata var. brevifolia A. Braun, 1863 (Sardegna)Isoetes velata var. tegulensis forma maculosa (Fiori) Fiori et Paol., Fl. Anal. Ital. 4: 10 (1907)Isoetes dubia var. maculosa Genn., Comment. Critt. Ital. l (2): 105 (1861); Isoetes velata var. tegulensis b. maculosa Genn., Ann. Agric. Sci. 175 (1865); Isoetes velata var. tegulensis forma emersa Fiori & Paol., Fl. Anal. Ital. 4: 10 (1907); Isoetes velata var. tegulensis c. emersa Genn., Giorn. Bot. Ital. 2: 96 (1870); Isoetes tiguliana Genn., Comment. Critt. Ital. 1 (1): 42 (1861)Isoetes velata calaostiensis Genn., Spec. e Var. Fl. Sarda, p. 31 (1867); Isoetes lacustris Moris, Stirp. Sard. EI. 3: 12 (1829); Isoetes velata forma sicula A. Br. ex Milde, Fil. Eur. & Atl., 282 (1867); Isoetes velata forma sicula Genn., Motel & Vendr., Actes Soc. Linn. Bord. 36: 384 (1883); Isoetes velata var. sicula Tod., Syn. Pl. Acot., 250 (1866); Isoetes velata var. typica b. sicula Ces. & DeNot.; Isoetes velata var. typica forma sicula Ces. & DeNot. apud Genn., Comment. Soc. Critt. Ital. 1 (2): 103. Sept (1861); Isoetes velata var. sicula Genn., Milde, Fil. Eur. & Atl. 282 (1867) - Algeria, Sardinia, Sicily. Isoetes velata subsp. tenuissima O. Bolós & Vigo, Butll. Inst. Catalana Hist. Nat. 38: 64 (1974) [Calamaria tenuissima (Boreau) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes tenuissima Boreau, Bull. Soc. Industr. Angers 21: 269 (1850), basion.; Isoetes tenuissima Mart., Cat. 341; Isoetes tenuissima var. tenuissima LeGrand, Bull. Soc. Bot. Fr. 42: 50 (1895); Isoetes velata var. tenuissima (Boreau) P. Fourn., 1934, Quatre Fl. Fr.: 5 (comb. illeg.); I. boryana subsp. tenuissima (Boreau) Douin in Bonnier, 1934, Fl. Compl. Fr., 12: 118 (comb. illeg.); Isoetes tenuissima var. capillifolia LeGrand, Bull. Soc. Bot. Fr. 42: 50 (1895); Isoetes tenuissima var. latifolia LeGrand, Bull. Soc. Bot. Fr. 42: 50 (1895); Isoetes tenuissima var. longifolia LeGrand, Bull. Soc. Bot. Fr. 42: 50 (1895); Isoetes velata subsp. tenuissima proles velata (A. Braun) Rouy, 1913, Fl. Fr., 14: 479 (comb. illeg.); Isoetes viollaei F. Hy, Bull. Herb. Boiss. Ill. App. 1: 23 (1894); Isoetes velata subsp. tenuissima [var.] viollaei (Hy) P. Fourn., 1934, Quatre Fl. Fr.: 5; Isoetes velata proles viollaei (Hy) Rouy, 1913, Fl. Fr., 14: 478; Isoetes chaboissaei Nyman, Consp. Fl. Eur., Suppl. 1: 871 (1883-84); Isoetes chaboissaei F. Hy, J. Bot. (Morot) 8: 97 (1894); Isoetes tenuissima var. chaboissaei LeGrand, Bull. Soc. Bot. Fr. 42: 50 (1895); Isoetes velata subsp. adspersa [var.] chaboissaei (Hy) P. Fourn., 1934, Quatre Fl. Fr.: 5] - Central France: Maine-et-Loire: étang de Beaurepaire (1988), Loire-et-Cher (1883), Indre (2003), Vienne (1895), Haute-Vienne (1947): étang de Ris-Chauvron (loc. class.), Creuse: étang de la Chaume (1985). Ill. BERTHET & LECOCQ, 1977: 349; Kasselmann: 73, 318. Isoetes velata subsp. boryana (Durieu) P. Fourn. (1934), Quatre Fl. Fr.: 5 (comb. illeg.); Isoetes boryana Durieu de Maisonneuve, 1861, Bull. Soc. Bot. France 8 (3): 164 = Calamaria boryana (Dur.) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Isoetes boryana brevis Gdgr., Fl. Eur. 27: 195 (1891); Isoetes boryana condensata Gdgr., Fl. Eur. 27: 195 (1891), Isoetes boryana legitima Gdgr., Fl. Eur. 27: 195 (1891); Isoetes boryana praetervisa Gdgr., Fl. Eur. 27: 195 (1891); Isoetes variabilis subsp. boryana (Durieu) P. Fourn., Quatre Fl. France: 5 (1934); Isoetes velata subsp. adspersa proles boryana (Durieu) Rouy, 1913, Fl. Fr., 14: 477 (comb. illeg.)] -  France: Gascony. Ill. C & K 1: 52, 59, Prelli: 83; BERTHET & LECOCQ, 1977: 351, 352. 

Isoetes vermiculata Hickey, 1989, Ann. Missouri Bot. Gard. 76(4): 1160-1161 - Venezuela: Depto. Atabapo: Tepui 2520-2560 m.

Isoetes viollaei F. Hy, 1894, J. Bot. 8: 97 (Etang de Ris-Chauvron) = I. velata susp. tenuissima

Isoetes virginica Pfeiffer = I. melanopoda

Isoetes weberi Herter ex Weber, 1922, Hedwigia 63: 255-256, fig. 44-45 - Uruguay, S Brazil: Rio Grande do Sul . Ill.  microspores  MUSSELMAN, 2002 (?= I. smithii)

Isoetes welwitschii A. Braun ex Kuhn, 1868, Filices Africanae,196 [Isoetes natalensis Baker, Handb. Fern Allies 132 (1887); Calamaria

 natalensis (Baker) Kuntze, Rev. Gen. Pl. 2: 828 (1891); Calamaria welwitschii (A. Braun ex Kuhn) Kunze, Revisio Generum Plantarum 2: 828 (1883)] - Sudan, Ethiopia, Eritrea, Ubangui,  Nigeria, Central African Republic; Ghana, Kenya, ZambiaAngola, Namibia, Transvaal, Natal,  Madagascar. Ill. COOK, 2004: 34. (this distribution is dependent on accepting the synonymy mentioned in the following note):

Annotations in the BM herbarium indicate that Jermy finally considered I. abyssinica to be conspecific with I. welwitschii A.Braun and also that I. nigritiana Kuhn and I. garnieri A.Chev. & P.Monnier should be included. I have not seen any of Chiovenda’s syntypes and it is possible they belong to more than one species. Pichi Sermolli refers to I. abyssinica in B.J.B.B. 53: 190 (1983) & 55: 197 (1985) but these Burundi specimens seem too robust as is also the case with material cited by Hall in Bot. J. Linn. Soc. 64: 117–139, fig. 2e, 3c, e, f, 4d, f, 5c, 6b, c (1971) from Ghana. Faden keeps up both species in U.K.W.F. ed. 2: 39 (1994) separating them as follows – megaspores with numerous large occasionally coalescing tubercles on the convex surface; triradiate facets completely covered by uniform tubercles I. welwitschii megaspores with fewer lower tubercles that never coalesce on the convex surface; triradiate facets with tubercles confined to centre or if more extensive then the ones in the centre the largest or occasionally tubercles lacking I. abyssinica He records I. welwitschii from Lolokwi (15 Apr. 1979, Gilbert 5366) and the S end of Mua Hills (2 Feb. 1969, Napper & Faden 1868). Burrows’ figures of the spore sculpture show very sparse verrucae quite unlike that of the type specimens.  Some specimens show characters which need investigation in the field to assess their possible importance. Several (e.g. Nairobi, 24 July 1977, Gilbert & Gillett 4800; Nairobi, Aug. 1977, Gilbert 4827; Nairobi National Park, 25 June 1971, Lye 6310) have dark marks on the outside of the sporangial area of the leaves (individual scattered dark epidermal cells) but initial suspicions this could be correlated with dark sporangia and velum rims ± 1 mm wide proved unfounded. Faden et al. 71/455 cited above has broadly triangular black corm scales with very cuspidate apices but few specimens have intact scales and the possible taxonomic significance of such characters can only be assessed in the field, but the fact that Gilbert & Mesfin 6500 (10 km Eldoret – Kitale, 8 Oct. 1981) is similar suggests it may be significant.Schippers records I. abyssinica from T 3 near Muheza and from along the coast (Wingfield 2032). The Muheza specimen has not been seen but the Wingfield specimen is far too large, particularly the leaf length and sporangia and is here referred to I. alstonii .Material from the Mathews Range also needs more study; Gilbert 5366 (Lolokwi [Ololokwe], 14 Apr. 1979) has sporangia 9 x 5 mm and the verrucae on large face of megaspores more oblong, slightly narrowed at the base. Gilbert 5002 cited above is not mature. (From aluka.org).

Isoetes wormaldii R. Sim, 1906, Trans. S. Afr. Phil. Soc. 16: 299, t. 5 - E Cape Prov. Ill. COOK, 2004: 34.

Isoetes yungiensis T.C. Huang & W. Carl Taylor, 2002, Novon 12 (4): 587-591 (2n = 22)- China: Yunnan: Guishou Plateau: Shashi Valley, Heilongtan, Tianshengqiao, Shuang Shao.

 

Hybrids

Isoetes x altonharvillii Musselman & R.D. Bray, 1995, Castanea 60(3): 246 - Virginia Remarks: Isoetes caroliniana x engelmannii

Isoetes x brittonii D.F.Brunt. & W.C.Taylor, 1990, American Fern Journal 80: 82-89 - Pennsylvania, Connecticut, New York = Isoetes engelmanii x Isoetes riparia

Isoetes x bruntonii Knepper & Musselman, 1996, Amer. Fern J. 86(1): 9 - Virginia = Isoetes engelmannii x hyemalis

Isoetes x carltaylorii Musselman L.J., Bray R.D., Knepper D.A., 1997, Canadian J. Bot. 75 (2): 301-309. = Isoetes acadiensis x Isoetes engelmannii

Isoetes x dodgei A.A. Eaton - North America = Isoetes setacea x Isoetes riparia

Isoetes x eatonii Dodge, 1897, Botanical Gazette 23: 16, t. 4-5, pro sp. = Isoetes setacea x Isoetes engelmannii [Isoetes eatoni Dodge, Ferns and Fern Allies N. Engl., 39 (1896); Isoetes eatoni forma gravesii (Eaton) Reed, Amer. Fern Journ. 35: 82 (1945); Isoetes eatoni var. gravesii (Eaton) Clute, Fern Bull. 16. 55 (1908); Isoetes x gravesii Eaton, Fernwort Papers 2: 14 (1900); Isoetes valida var. gravesii (Eaton) Clute, Fern Allies, 243 (1905)] - New Hampshire, Massachusetts, New Jersey, Connecticut. Ill. FASSETT: 46, map Muenscher: 342.

Isoetes setacea x riparia Britton, D.M. & D.F. Brunton, 1989. A new Isoetes hybrid (Isoetes setacea x riparia) for Canada. Canad. J. Bot. 67 (10): 2995-3002 - illus., maps.

Isoetes x echtuckeri D.F.Brunton & D.M.Britton, 1999, hybr. nov., a new triploid quillwort from northeastern North America. Canad. J. Bot. 77 (11): 1662-1668 - illus.

Isoetes x fairbrothersii J.D.Montgom. & W.C.Taylor, 1994, Amer. Fern J. 84(4): 120 (1994) - New Jersey = Isoetes engelmannii x lacustris

Isoetes x foveolata A.A. Eaton ex Dodge, Ferns & Fern Allies of New England 38 (1896) - North America = Isoetes engelmannii x Isoetes tuckermanii

Isoetes x harveyi A.A. Eaton - North America = Isoetes lacustris x Isoetes tuckermanii (Isoetes tuckermannii var. harveyi (Eaton) Clute

Isoetes x altonharvillii L.J. Musselman & R.D. Bray = Isoetes caroliniana x Isoetes engelmannii

Isoetes x heterospora A.A. Eaton = I. lacustris x Isoetes tuckermannii (I. tuckermannii var. heterospora (Eaton) Clute

Isoetes x hickeyi W.C.Taylor & Luebke, 1988, Amer. Fern J. 78(1): 6 - Wisconsin = Isoetes setacea x Isoetes lacustris

Isoetes x jeffreyi D.M. Britton & D.F.Brunton, 1992, Canadian Journal of Botany 70: 447-452 - Quebec = Isoetes lacustris x Isoetes riparia, Isoetes macrospora x riparia

Isoetes x marensis D.M.Britton & D.F.Brunton, 1995, Canad. J. Bot. 73 (9): 1352 - British Colombia = Isoetes howellii x maritima

Isoetes x michinokuana M. Takamiya & al. - Distribution unknown

Isoetes x pseudotruncata D.M.Britton & D.F.Brunton, 1996, Canad. J. Bot. 74(1): 58 - British Colombia = Isoetes setacea x maritima

Isoetes x truncata (A.A. Eaton) Clute, 1905, The Fern Allies 260, SE Alaska [Isoetes maritima x Isoetes occidentalis = Isoetes echinospora var. truncata A.A. Eaton, Working List of North American Pteridophytes 27 (1901) = Isoetes muricata subsp. truncata (A.A.Eaton) H.P.Fuchs, Proc. Kon. Ned. Akad. Wetensch. C 85(2): 239 (1982)].